5.GENERAL KEY No 2 to PTEROPSIDA

1Aquatic plants.
1'Land plants or epiphytes.
2Plants floating; leaves small, simple or bilobed
2'Plants rooted in earth or on rocks; leaves larger, more divided.
3Leaves 4-partite; sporocarps attached to stipes
3'Leaves not 4-partite; sporangia singly or in sori on lower surface of lamina.
4Sporangia borne singly, protected by reflexed edges of narrow lamina.
4'Sporangia grouped in sori, on lower surface of lamina, not protected by reflexed edges.
5Fern of stream-beds in deep shade; fronds pinnatifid, sori without indusia
5'Fern of open swamps; fronds bipinnatifid, sori indusiate.
6Stipe containing 3 or more vascular bundles throughout.
6'Stipe containing one vascular bundle, or two at the base joining upwards to form a single Strand of various shape.
7Rhizome in most cases dorsiventral, stipes jointed to it or to phyllopodia borne by rhizome.
7'Rhizome not dorsiventral, or if so the stipes not jointed to it.
8Sori of various form (in some cases acrostichoid), not indusiate.
8'Sori indusiate.
9Fronds pinnate; pinnae articulate to rachis, veins free
9'Fronds rarely pinnate; pinnae (if present) not articulate to rachis, or if so veins reticulate.
10Acrostichoid; spores with perispore; veins free or uniting near margin
10'Acrostichoid or not; no perispore; where acrostichoid, veins much anastomosing.
11Fronds simple.
11'Fronds more or less branched.
12Sori near midrib of frond
12'Sori at ends of veins, solitary or joining many veins
13Pinnae jointed to rachis
13'Pinnae not jointed to rachis
14Stipes containing numerous bundles in a complex pattern, not a simple ring, in cross section.
14'Stipes containing a simple open ring of bundles.
15Rootstock massive, short or tall; if creeping, subterranean.
15'Rootstock slender, long-creeping or high-climbing
16Mangrove plants; distal pinnae acrostichoid, lower ones sterile.
16'Not mangrove plants; not acrostichoid.
17Stock and stipes containing woody tissue; no stipules at bases of stipes.
17'Stock and stipes without woody tissue; stipules present at bases of stipes.
18Veins free
18'Veins anastomosing
19Sporangia on a separate branch attached at base of lamina, or to surface of lamina; no annulus.
19'Sporangia not on a separate branch of frond; annulus present.
20Fronds pinnate or bipinnate, leaflets jointed to rachis.
20'Leaflets (if present) not jointed to rachis.
21Fronds in close tufts, not spaced on a climbing rhizome; sori separate and indusiate.
21'Fronds spaced on a climbing rhizome; sori not indusiate, in most cases acrostichoid.
22Fronds simple, sori oblique, exindusiate
22'Fronds otherwise.
23Sori elongate, either close to midrib or to edge.
23'Sori not elongate near midrib or edge.
24Sori close to midrib
24'Sori close to edge
25Sporangia acrostichoid, edge of lamina in some cases reflexed.
25'Sporangia not acrostichoid.
26Veins of sterile leaflets free.
26'Veins of sterile leaflets anastomosing.
27Fronds bipinnate
27'Fronds simply pinnate.
28Rhizome creeping, dorsiventral
28'Rhizome erect, radially arranged
29Rhizome creeping, dorsiventral
29'Rhizome erect, not dorsiventral
30Rachis grooved, groove open to admit branches or midribs of leaflets.
30'Rachis not grooved, its wings (if any) confluent with edges of lamina.
31Rhizome-apex and bases of stipes hairy or apparently glabrous, not scaly.
31'Rhizome-apex and bases of stipes scaly.
32Fronds repeatedly pseudo-dichotomous, a dormant apex between pairs of branches.
32'Fronds otherwise branched.
33Apices of some or all primary rachis-branches dormant.
33'Apices of primary rachis-branches not dormant (apex of main rachis sometimes periodically dormant).
34Rachis climbing and twining; sporangia solitary
34'Rachis trailing, not twining; several sporangia in a sorus, indusiate.
35Sori at ends of veins, either in cups or protected by two equal or subequal flaps.
35'Sori otherwise.
36Filmy ferns; lamina one cell thick
36'Not filmy, lamina thicker.
37Massive rootstock, prostrate or erect and trunk-like, densely covered with long hairs.
37'Slender creeping rhizome; hairs short
38Bases of stipes bearing prominent aerophores; fronds dimorphous, fertile with narrow acrostichoid pinnae
38'Bases of stipes lacking aerophores; fertile fronds otherwise.
39Sporangia large, round, lacking an annulus, on deeply dissected pinnae
39'Sporangia otherwise.
40Sporangia variously arranged on lower surface of fronds, usually along the veins, or in a submarginal groove, not indusiate.
40'Sporangia arranged otherwise.
41Sori elongate
41'Sori small and round, or acrostichoid
42Annulus almost apical; fronds simple or branched dichotomously; sporangia on special appendages
42'Annulus and frond otherwise.
43Fronds pedately branched; sori superficial, indusiate
43'Fronds pinnately branched; sori at ends of veins, indusiate or not.
44Sporangia on surface of reflexed marginal lobes
44'Sporangia not on such lobes.
45Sori elongate along the veins of the lower surface.
45'Sori not elongate along the veins of the lower surface (in some cases elongate along a submarginal vein).
46Sori indusiate.
46'Sori not indusiate.
47Scales clathrate; the 2 vascular bundles at base of stipe uniting upwards to an X-shape.
47'Scales not clathrate; the 2 vascular bundles at base of stipe uniting upwards to a U-shape.
48Unicellular slender hairs lacking
48'Unicellular slender hairs on frond and on scales
49Fronds simple; sori in grooves (or rarely superficial); epiphytes or rock-plants.
49'Fronds usually not simple; sori not in grooves; not epiphytes or rock-plants.
50Sori not occupying whole length of veins.
50'Sori occupying whole length of veins.
51Lower surface of lamina not covered with powder
51'Lower surface of lamina covered with powder
52Slender unicellular hairs lacking on lamina and scales.
52'Slender unicellular hairs present on lamina and scales.
53Sori in marginal or submarginal grooves, or superficial and forming continuous lines parallel to midrib.
53'Sori otherwise.
54Scales clathrate
54'Scales not clathrate
55Sori at ends of veins (uniting several veins or not) protected by reflexed edge of lamina.
55'Sori not at ends of veins or not so protected.
56Rachis grooved on upper surface, groove opening to admit groove of midrib of pinna.
56'Rachis not grooved in this way
57Lower surface of frond more or less covered with white or yellow powder.
57'Lower surface of frond not so covered.
58Sori submarginal, at ends of veins, in small projecting cups, or protected by an indusium attached on the side remote from the margin.
58'Sori otherwise.
59Sori in cups projecting from margin
59'Sori not in such cups.
60Sori always solitary at ends of Single veins; scales of rhizome bearing papillae on marginal cells; rhizome not protostelic
60'Sori often uniting ends of several veins; scales lacking papillae; rhizome protostelic.
61Main rachis of fronds periodic in growth; thicket-forming ferns.
61'Main rachis of fronds continuous in growth; not thicket-forming ferns.
62Small epiphytes (occasionally on rocks); sori superficial or immersed in pockets in substance of the lamina, not indusiate
62'Not epiphytes; sori in most cases indusiate.
63Hairs on fronds unicellular (in rare cases very long hairs may consist of more than one cell).
63'Hairs on fronds multicellular, cells short
64Scales confined to swollen basal part of stipe; no superficial hairs or glands on scales.
64'Scales not so confined, bearing superficial unicellular hairs or glands.