KEY TO THE SPECIES

1Leaves 1-foliolate (or rarely 3-foliolate).
1'Leaves 1-4-pinnate.
2Flowers 4-merous. Fruit usually 4-seeded. Philippines.
2'Flowers 5-merous. Fruits usually 6-seeded (sometimes less by abortion). Not in Philippines.
3Leaflets obovate, pair of foliar outgrowths (rarely seen as reduced leaflets) above the stipular wing
3'Leaflets elliptic or ovate, foliar outgrowths absent.
4Seeds complexely ruminate (Fig. 4a). Leaflets up to 22 by 9 cm
4'Seeds simply ruminate (compare Fig. 4d-e). Leaflets usually over 22 by 9 cm.
5Corolla tube + staminodial lobes less than 3 mm long, lower free part of staminodial tube up to ½ mm. Fruit usually up to 10 mm ø Malaya, Sumatra, Java
5'Corolla tube + staminodial lobes over 3 mm long, lower free part of staminodial tube over ½ mm. Fruit usually over 10 mm ø New Guinea.
6Corolla tube + staminodial lobes less than 4½ mm. Leaflet base subauriculate.
6'Corolla tube + staminodial lobes over 4½ mm. Leaflet base cuneate to truncate
7Stipule a narrow wing, somewhat persistent, scar long and thin. Compare Fig. 2(4, 7, 13).
7'Stipule obovate, caducous, leaving a broad scar. Compare Fig. 3(21, 23, 24).
8Flowers 4-merous. Fruits usually 4-seeded (if 6-seeded then fruit over 20 mm ø).
8'Flowers 5-merous. Fruit usually 6-seeded.
9Staminodial tube over 5 mm long. Fruits over 20 mm ø, 6-seeded. New Guinea and Solomon Is.
9'Staminodial tube less than 5 mm. Fruit less than 20 mm ø, 4-seeded. Philippines.
10Lobes of staminodial tube strongly bifid. Inflorescences condensed, erect. Flowers orange-yellow. Mainland of New Guinea
10'Lobes of staminodial tube retuse. Inflorescences lax, pendulous. Flowers white. Solomon Is.
11Style over 3 mm long; staminodial tube 4-5 mm long; filaments over 2 mm. Inflorescence generally condensed, 3-branched, peduncle usually up to 3 cm. Young parts sometimes fulvously pubescent. Leaflets usually elliptic to elliptic-lanceolate, over 7 cm wide; ultimate venation distinct. Rumination outline in seed simple to slightly branched
11'Style up to 2 mm long; staminodial tube 2½-4 mm; filaments up to 2 mm. Inflorescence generally lax and multi-branched, peduncle over 3 cm. Young parts never fulvous pubescent. Leaflets generally ovate to ovate-lanceolate and less than 7 cm wide; ultimate venation indistinct. Rumination outline in seed complexly reticulate
12Stems spiny.
12'Stems not spiny or information lacking.
13Leaves 1-pinnate
13'Leaves 2- or 3-4-pinnate
14Corolla tube + staminodial lobes 6 mm or more and staminodial tube over 5¼ mm long. Fruit where known over 20 mm ø. New Guinea and Solomon Is.
14'Corolla tube + staminodial lobes up to 6 mm or staminodial tube less than 5¼ mm long. Fruit less than 20 mm ø. Not in New Guinea except for L. gonioptera and L. aculeata.
15Leaves 3- or 4-pinnate, leaflets up to 14 by 5 cm. Flowers pink. Mainland New Guinea
15'Leaves 1- or 2-pinnate, leaflets mostly over 14 by 5 cm. Flowers not pink.
16Lobes of staminodial tube strongly bifid. Inflorescence condensed, erect. Flowers orange-yellow. Mainland New Guinea
16'Lobes of staminodial tube retuse. Inflorescence lax, pendulous. Bismarck Archipelago and Solomon Is.
17Inflorescence usually glabrous. Corolla tube + staminodial lobes 8-11 mm, filaments 5-7 mm, anthers 3-5 mm. Young parts not fulvously pubescent. Bismarck Archipelago
17'Inflorescence usually fulvously pubescent. Corolla tube + staminodial lobes 6-8 mm, filaments 3 mm, anthers 2 mm. Young parts usually fulvously pubescent. Solomon Is.
18Leaves 1-pinnate.
18'Leaves 2- to 4-pinnate.
19Corolla tube + staminodial lobes up to 2½ mm, staminodial tube up to 1½ mm long. Small shrubs with creeping rootstock
19'Corolla tube + staminodial lobes over 2½ mm; staminodial tube over 1½ mm. Shrubs or trees.
20Flowers red
20'Flowers green or white.
21Calyx ± inflated around the corolla tube, completely enclosing the corolla in the bud. Corolla tube + staminodial lobes over 5 mm, staminodial tube over 4 mm. Fruit c. 20 mm ø
21'Calyx not enclosing the corolla in the bud, not so inflated. Corolla tube + staminodial lobes less than 5 mm, staminodial tube less than 4 mm. Fruit usually less than 15 mm (rarely to 20 mm) ø.
22Leaflet base subauriculate. Stem smooth. Staminodial tube up to 3 mm long, lower free part up to 1¼ mm long. New Guinea
22'Leaflet base rounded to cuneate. Stem spiny. Staminodial tube 3-3½ mm long, lower free part over 1¼ mm long. Scattered throughout Malesia, except Malaya, very rare in W. New Guinea
23Staminodial tube up to 2¼ mm long. Stipules up to 6 cm long. Petiole generally less than 10 cm.
24Corolla tube + staminodial lobes over 3¼ mm, sinuses of staminodial tube shallow. Flowers greenish white. Fruit greyish blue. Stems and ultimate branches spiny.
24'Corolla tube + staminodial lobes at least 2¼ mm long, sinuses of staminodial tube shallow. Flowers red. Fruit red. Stems and branches not spiny
24''Staminodial tube at least 2½ mm long. Stipules usually over 6 cm long. Petiole generally over 10 cm
25Flowers 4-merous. Fruits usually 4-seeded. Philippines
25'Flowers 5-merous. Fruits usually 6-seeded.
26Leaves 1-pinnate.
26'Leaves 2-4-pinnate.
27Flowers red to orange-yellow
27'Flowers greenish white.
28Corolla tube + staminodial lobes over 4 mm, staminodial tube over 2¾ mm. Fruit c. 20 mm ø.
28'Corolla tube + staminodial lobes less than 4 mm, staminodial tube less than 2¾ mm. Fruit up to 15 mm ø
29Sinuses of staminodial tube deep, c. 1 mm. Leaflets generally over 30 cm long.
29'Sinuses of staminodial tube shallow to ½ mm. Leaflets generally but not exclusively up to 30 cm long
30Corolla tube + staminodial lobes over 4 mm, staminodial tube over 3 mm. Fruit at least 20 mm ø. New Guinea.
30'Corolla tube + staminodial lobes less than 4 mm, staminodial tube less than 3 mm. Fruit less than 20 mm ø.
31Sinuses of staminodial tube deep, c. 1 mm. Leaflets generally over 30 by 9 cm, nerves usually over 10 pairs. Stipular scar generally over 4 cm long
31'Sinuses of staminodial tube shallow, less than ½ mm. Leaflets generally less than 30 by 9 cm, nerves generally less than 10 pairs. Stipular scar up to 4 cm long.
32Inflorescence and leaflets with large, discoidal, pallid brown pearl glands. Bracts of inflorescence large, conspicuous, up to 8 by 5 mm
32'Pearl glands inconspicuous, or absent. Bracts smaller, inconspicuous.
33Petiole, rachis, and costa with crisped fluted emergences. Leaflets large, 30-40 by 10-17 cm. Celebes
33'Emergences absent. Leaflets usually, but not exclusively, smaller.
34Flowers greenish-white
34'Flowers red to orange-yellow.
35Corolla tube + staminodial lobes generally over 3 mm, staminodial tube over 2 mm, sinuses of staminodial tube shallow
35'Corolla tube + staminodial lobes up to 3 mm, staminodial tube less than 2 mm, sinuses of staminodial tube deep