Malesia: Sumatra, Peninsular Thailand, Peninsular Malaysia, Java, Borneo, Celebes, New Guinea.
Typical specimens have deeply dissected fertile pinnae, with sori nearly marginal, on separate teeth, and the lamina between the sori cut to c. ½. However, many specimens (especially from the eastern part of the range) have less deeply divided pinnae, and such specimens can easily be confused with other species. They can best be distinguished by the rhachis indument, which consists of scales with a narrow, dark acumen, sometimes needle-like, often directed sideways towards the upper surface. Hairs are sometimes present on the upper surface of the costae, usually near the base only.