KEY TO THE SPECIES

1Small scales on frond stellate or elongate, their marginal hairs glandularor with a glandularapical cell, thick-walled unicellular marginal hairs lacking; rhizome-scales various.
1'Small scales on frond (except near edge) ± orbicular, pale, with marginal teeth or hairs each of one thick-walled cell, not glandular; rhizome-scales always narrow, rigid, dark, glossy.
2Lamina of sterile fronds rarely over 7 cm long; rhizome 1-2 mm ø, fronds usually well-spaced on it.
2'Lamina of sterile fronds usually more than 7 cm long; rhizome thicker.
3Rhizome-scales light brown.
3'Rhizome-scales dark
4Sterile lamina elliptic, commonly 4-7 cm long, base decurrent as a narrow wing.
4'Sterile lamina ovate, almost same shape and size as fertile, c. 2 cm long.
5Fertile lamina spathulate, abruptly contracted at base and then decurrent as a narrow wing 2 cm long; rhizome-scales to 1 mm wide
5'Fertile lamina narrowly elliptic, little decurrent; rhizome-scales 2 mm wide
6Rhizome long-creeping with well-spaced fronds.
6'Rhizome short, with closely-placed fronds.
7Apex of sterile fronds acute or short-acuminate.
7'Apex of sterile frond rounded.
8Scales dark, glossy, 1 mm wide
8'Scales thin, medium brown, 2-3 mm wide
9Rhizome-scales light brown; scales on frond red-brown
9'Rhizome-scales dark brown; scales on frond dark, glossy (not known in no 9).
10Rhizome-scales to 5 by 1.5 mm; stipe of sterile fronds 4-15 cm long.
10'Rhizome-scales shorter, less than 1 mm wide; stipe of sterile fronds 3-5 cm.
11Lamina of sterile frond decurrent to joint with phyllopodium or within 2 cm of joint.
11'Lamina of sterile frond not thus decurrent.
12Edge of sterile lamina conspicuously pale and thin, 0.5 mm or more wide.
12'Edge of sterile lamina not conspicuously pale and thin.
13Sterile lamina twice as wide as fertile.
13'Sterile lamina little wider than fertile
14Apex of sterile frond bluntly pointed or rounded, base cuneate and then decurrent for 4-7 cm as a wing 1-2 mm wide on each side of costa
14'Apex of sterile frond short-acuminate, base narrowed evenly almost to joint with phyllopodium
15Sterile frond acuminate or apiculate.
15'Sterile frond not acuminate nor apiculate.
16Rhizome-scales 10-15 by 3-4 mm, light brown, thin; lamina widest above middle
16'Rhizome-scales much smaller, dark; lamina widest at middle.
17Sterile frond c. 22 by 2 cm
17'Sterile frond much larger, to 4½ cm wide.
18Apex of sterile frond acuminate.
18'Apex of sterile frond broad with a narrow apiculus 5 mm long.
19Sterile frond 35-50 by 3.5-5 cm
19'Sterile frond to 35 by 2.3 cm
20Sterile frond widest 1/3 from apex, narrowed rather abruptly to broad-pointed apex.
20'Sterile frond widest about middle, or if above middle narrowed gradually to a rounded apex.
21Rhizome-scales 15-20 mm long, 2 mm wide; fronds thick, veins obscure; costa broad and little prominent on lower surface
21'Rhizome-scales not over 10 mm long; fronds rather thin, veins distinct; costa slender and prominent on lower surface.
22Scales on surface of frond pale, bearing slender hairs 1 mm long; stipe of fertile frond 10-15 cm
22'Scales on frond dark, bearing very short hairs; stipe of fertile frond 25-30 cm
23Sterile frond widest at middle, apex acute; fertile frond to 2 cm wide
23'Sterile frond widest a little above middle, gradually narrowed to rounded apex; fertile fronds not over 1.3 cm wide
24Scales on rhizome dark, rigid, glossy.
24'Scales on rhizome medium to light brown, mostly rather thin.
25Small scales on upper surface of frond dark, glossy, elongate, with dark rigid cylindrical marginal hairs each consisting of one cell with a small thin-walled cell at its apex.
25'Small scales on upper surface otherwise, at least some of their marginal hairs multicellular with glandular apical cell.
26Scales on lamina 1-2 mm long; fronds with broadly rounded apex.
26'Scales on lamina much smaller; apex of fronds not broadly rounded.
27Apex of sterile frond rounded.
27'Apex of sterile fronds acute or acuminate.
28Phyllopodia ½-1 cm long; lamina of sterile frond always c. 10 times as long as wide.
28'Phyllopodia 1.5-2 cm long; lamina of sterile frond often proportionately broader
29Scales on costa and stipe appressed, on lamina very small
29'Scales on stipe and costa spreading, on lamina 1-2 mm long, abundant
30Lamina of sterile frond c. 10 times as long as wide.
30'Lamina of sterile frond c. 5 times as long as wide.
31Rhizome-scales less than 1 mm wide, somewhat crisped; stipe of fertile frond not or little longer than that of sterile
31'Rhizome-scales more than 1 mm wide, flat; stipe of fertile frond usually twice as long as sterile.
32Sterile frond 3.5-5 cm wide, 35-50 cm long
32'Sterile frond to 2.3 cm wide, 35 cm long
33Rhizome bearing fronds 5-10 mm apart; scales 6 by 1 mm; sterile lamina widest below middle, rather abruptly contracted at base, apex short-pointed.
33'Rhizome bearing fronds close together; scales 10 by 1.5 mm; sterile lamina widest at middle, base rather narrowly cuneate, apex acuminate
34Lower surface of sterile lamina covered with a feit of narrow scales 2-3 mm long
34'Lower surface of lamina bearing smaller, usually appressed, scales.
35Fronds of adult plants widest above middle, with rounded apices and decurrent bases.
35'Fronds of adult plants, if widest above middle, rather much narrowed upwards, apex not broadly rounded.
36Fronds lacking a fringe of spreading scales.
36'Fronds bearing a fringe of spreading scales
37Rhizome-scales less than 1 mm wide
37'Rhizome-scales 2 mm or more wide
38Sterile fronds with distinct thin pale edge which bears abundant scales 1-1.5 mm long, persistent on young plants, mostly lost on old ones
38'Sterile fronds without a fringe of scales.
39Sterile lamina twice as long as wide (to 12 by 6 cm); stipe of fertile frond as long as sterile frond with its stipe
39'Not this combination of characters.
40Apex of sterile frond distinctly pointed, point broad or acute.
40'Apex of sterile frond distinctly rounded.
41Sterile frond widest in distal half, base very gradually and evenly narrowed, stipe to 5 cm long
41'Sterile frond widest at or below middle, base otherwise, stipe in most cases longer.
42Pale margin of sterile frond thin, flat, 0.5 mm wide or nearly so.
42'Pale margin of sterile frond very narrow, usually reflexed.
43Fronds widest below middle, narrowed gradually to apex; veins distinct
43'Fronds widest at middle, apex obtuse-angled; veins not distinct
44Scales to 1.5 mm wide, not flat.
44'Scales wider, flat or little crisped, edges not inrolled.
45Scales straight and rather stiff with inrolled edges; frond commonly 35 cm or more long, narrowly elliptic
45'Scales thin, crisped throughout, with long twisted hair-points; frond rarely 30 cm long, broadly elliptic
46Stipe of fertile frond not much longer than stipe of sterile.
46'Stipe of fertile frond at least twice as long as stipe of sterile.
47Scales commonly 20-25 by 2.5-3 mm
47'Scales not over 10 mm long.
48Rhizome-scales with many spreading marginal hairs; scales on lamina to 1 mm long, narrow
48'Rhizome-scales with few marginal hairs; scales on lamina stellate, under ½ mm long.
49Costa broad and not prominent beneath; rhizome-scales firm
49'Costa prominent beneath; rhizome-scales thin, with isodiametric cells
50Scales 6 by 2.5 mm, sterile frond short-acuminate.
50'Scales to 15 by 2 mm; sterile frond broadly pointed
51Rhizome-scales 3-5 mm long; fertile frond c. 8 by 3.3 cm
51'Rhizome-scales 10-15 mm long; fertile frond at least 6 times as long as wide.
52Spores 52-59 μm long with broad perispore of few folds; stipe of fertile frond twice as long as that of sterile; edge of lamina thick, deflexed; lamina greenish when dry.
52'Spores c. 37 μm long with narrow much-folded perispore; stipe of fertile fronds not much longer than of sterile; pale edge of lamina thin; fronds brownish when dry, thick, rigid
53Lamina of sterile frond to 30 by 5 cm, stipe 5-8 cm long.
53'Lamina of sterile frond 35-55 cm long, stipe 14-18 cm long.
54Scales on upper surface conspicuously stellate; sterile fronds often to 3 cm wide.
54'Scales on upper surface with very short marginal hairs; sterile fronds c. 1 cm wide
55Scales on lower surface very small, each with 3-5 marginal hairs ½-1 mm long spreading away from frond-surface
55'Scales on lower surface with a distinct lamina (flat or not) bearing more slender and shorter marginal hairs.
56Scales of lower surface mostly elongate, bullate at base; on upper surface flat, those near margin elongate, forming a conspicuous spreading fringe
56'Scales of lower surface small, orbicular, flat or cup-shaped; scales of upper surface flat, about same size as on lower, those near margin not forming a conspicuous fringe