Drynaria

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Drynaria

Description

Epiphytic, epilithic or terrestrial.

Distribution

Asia-Tropical: Paleotropical: present
Paleotropical. Throughout Malesia.

Morphology

When the base fronds disintegrate, some species retain the naked rachises for a long time, sometimes with an empty mesh of finer veins.

Anatomy

Anatomy:

Taxonomy

Drynaria is a distinct genus, best recognisable by the peculiar frond dimorphism between sterile base fronds and fertile foliage fronds, the distinctive leathery texture of the lamina, a distinctive venation pattern and the presence of nectaries near the bases of the pinnae. The last three characters are shared with Aglaomorpha. A curious character unique to Drynaria is the aborted apex, with a lateral pinna taking its place, giving a peculiar ‘lop-sided’ look to the apical part of most fronds.

Together with Aglaomorpha, Drynaria forms a distinct group, recognised as subfamily Drynarioideae by Crabbe, Jermy & Thomas (), and as tribe Drynarieae by Hennipman et al. (1990). Roos (1985) presents a cladistic analysis but does not propose a subdivision of Drynaria based on his analysis.

Citation

in Sp. Fil. 1864: 93
Copel., Gen. Fil. 1947: 204
Hook. & Bauer, Gen. Fil. 1842: t. 21
Alderw., Malayan Ferns. 1917: 415
Tind., Contr. NSW Nat. Herb. 1961: 29
Ching - in Sunyatsenia. 1940: 261
Hennipman et al. - in Kramer & Green, Fam. & Genera Vasc. Pl. 1. 1990: 212
Holttum - in Revis. Fl. Malaya. 1954: 182
Copel., Polypod. Philipp. 1905: 135
Bedd., Handb. Ferns Brit. India. 1883: 338
Baker, Syn. Fil. 1868: 366
Hook., Gard. Ferns. 1862: pl. 1
Copel. - in Univ. Calif. Publ. Bot. 1929: 117
Copel., Gen. Fil. 1947: 203
Alderw., Malayan Ferns. 1908: 696
J. Sm., Hist. Fil. 1875: 107
Alderw., Malayan Ferns. 1917: 415
Fée, Gen. Filic. 1850-1852: 269
Copel. - in Philipp. J. Sc. Bot. 1911: 140
Morton - in Taxon. 1970: 647
Alderw., Malayan Ferns. 1908: 697
Copel., Fern Fl. Philipp. 1960: 496
Pichi Serm. - in Webbia. 1977: 379
Backer & Posth., Varenfl. Java. 1939: 232
Roos, Drynarioideae. 1985: 255
Alderw., Malayan Ferns. 1908: 679
Brownlie - in Fl. Nouv. Caléd. et Dép. 1969: 278
Bedd., Ferns S. India. 1863: 63
Malayan Ferns. 1917: 415
Diels - in Bot. Jahrb. Syst. 1900: 207
Tardieu & C. Chr. - in Fl. Indo-Chine. 1941: 516
J. Sm., Ferns Brit. & For. 1866: 102