Bolbitis heteroclita

Distribution

Asia-Tropical: Caroline Is: present E. India: present Lesser Sunda Is: absent Melanesia: present Micronesia: present S. Japan: present Solomon Is: present Southern America:, Argentina Northeast (Formosa present)
E. India to S. Japan, Formosa, and Indo-China; in Malesia: throughout, except for the Lesser Sunda Is.; also in Micronesia (Caroline Is.) and Melanesia (Solomon Is.).

Notes

The morphological variation is outstanding. Throughout its area forms with simple fronds occur, e.g. B. simplicifolia from higher elevations in Malaya. In the Philippines several distinct vegetatively propagating dwarfs occur, e.g. Edanyoa difformis and B. cuspidata. Autoploidisa- tion and hybridisation are common. Field studies are necessary to further unravel this aggregate.

Citation

Backer & Posth., Varenfl. Java. 1939: 80
Bl., En. Pl. Jav. 1828: 104
v.A.v.R. - in Handb. Mal. Ferns. 1917: 436
Copel., Fern Fl. Philip. 1960: 261
Copel. - in Philip. J. Sc. 1928: 393
Ching, Fl. Hain. 1964: f. 75
v.A.v.R. - in Handb. Mal. Ferns. 1917: 60
C. Chr. - in Copel., Philip. J. Sc. 37. 1928: 411
Copel., Fern Fl. Philip. 1960: 258
Backer & Posth. - in Nat. Tijd. N. I. 1933: 163
v.A.v.R., Handb. Mal. Ferns. 1908: 735
Hook. & Grev., Fl. Java Fil. 1829: pl. 13
v.A.v.R., Handb. Mal. Ferns. 1908: 739
Copel., Fern Fl. Philip. 1960: 261
Copel., Fern Fl. Philip. 1960: 259
Backer & Posth. - in Nat. Tijd. N. I. 1933: 159
Copel., Fern Fl. Philip. 1960: 265
Copel., Polyp. Philip. 1905: 42
Hennipman - in Leid. Bot. Ser. 1977: f. 60
Copel., Polyp. Philip. 1905: 4
Holttum, Ferns Malaya. 1954: f. 272
Copel., Polyp. Philip. 1905: 43
Copel., Fern Fl. Philip. 1960: 258
Copel., Fern Fl. Philip. 1960: 257
Holttum, Ferns Malaya. 1954: f. 271
Backer & Posth., Varenfl. Java. 1939: f. 13
Fee, Suppl. Corr. 1917: 60
DeVol & Kuo - in Fl. Taiwan. 1975: pl. 122

Synonymy

Bolbitis heteroclita (Presl) Ching in C. Chr., Ind. Fil.: 48. 1934
  • Acrostichum heteroclitum Presl, Rel. Haenk.: pl. 2: f. 2. 1825
  • Heteroneuron heteroclitum Fee, Hist. Acrost.: 92. 1845
  • Chrysodium heteroclitum Kuhn in Ann. Mus. Bot. Lugd.-Bat. 4: 294. 1869
  • Leptochilus heteroclitus C. Chr., Ind. Fil.: 385. 1906
  • Campium heteroclitum Copel. in Philip. J. Sc. 37: f. 13. 1928
  • =Nephrodium cuspidatum Presl, Rel. Haenk.: 31. 1825
  • Polystichum cuspidatum Presl, Tent. Pterid.: 82. 1836
  • Heteroneuron cuspidatum Presl, Epim. Bot.: 169. 1851
  • Chrysodium cuspidatum Kuhn in Ann. Mus. Bot. Lugd.-Bat. 4: 293. 1869
  • Leptochilus cuspidatus C. Chr., Ind. Fil.: 384. 1906
  • Campium cuspidatum Copel. in Philip. J. Sc. 37: f. 19. 1928
  • B. cuspidata Ching in C. Chr., Ind. Fil.: 48. 1934
  • =Acrostichum flagelliferum Wall. ex Hook. & Grev., Ic. Fil.: pl. 23. 1827
  • B. flagellifera Schott, Gen. Fil.: adt. 13. 1835
  • Gymnopteris flagellifera Bedd., Ferns Br. India: 27. 1876
  • =Acrostichum proliferum Bl., En. Pl. Jav.: 104. 1828
  • =Leptochilus linnaeanus Fee, Hist. Acrost.: pl. 47: f. 2. 1845
  • Dendroglossa linnaeana Fee, Gen. Polyp.: 81. 1852
  • Gymnopteris linnaeana Christ in J. de Bot. 19: 125. 1905
  • Leptochilus heteroclitus var. linnaeanus Christ in Philip. J. Sc. 2: Bot. 160. 1907
  • Campium linnaeanum Copel. in Philip. J. Sc. 37: 343. 1928
  • B. linnaeana C. Chr., Ind. Fil.: 198. 1934
  • =Cyrtogonium acuminatum Brackenr. in Wilkes, U.S. Expl. Exp. 16: 86. 1854
  • =Poecilopteris stenophylla Kurz ex T. & B. in Nat. Tijd. N. I. 27: 15. 1864
  • B. stenophylla Ching in C. Chr., Ind. Fil.: 50. 1934
  • =Chrysodium heteroclitum var. subcrenatum Kuhn in Ann. Mus. Bot. Lugd.-Bat. 4: 294. 1869
  • =Acrostichum modestum Baker in J. Linn. Soc. 22: 231. 1886
  • Leptochilus modestus C. Chr., Ind. Fil.: 386. 1906
  • Campium modestum Copel. in Brittonia 1: f. 1. 1931
  • B. modesta Ching in C. Chr., Ind. Fil.: 49. 1934
  • =Gymnopteris inconstans Copel. in Perk., Fragm. Fl. Philip.: 177. 1905
  • Leptochilus inconstans Christ in Bull. Herb. Boiss. II: 1005. 1906
  • Leptochilus heteroclitus var. inconstans Christ in Philip. J. Sc. 2: Bot. 160. 1907
  • B. inconstans Ching in C. Chr., Ind. Fil.: 48. 1934
  • =Leptochilus heteroclitus var. euryhasis Christ in Philip. J. Sc. 2: Bot. 159. 1907
  • Campium heteroclitum var. eurybasis Copel. in Philip. J. Sc. 37: 361. 1928
  • =Leptochilus heteroclitus var. foxworthyi Christ in Philip. J. Sc. 2: Bot. 160. 1907
  • =Leptochilus sumatranus v.A.v.R. in Bull. Jard. Bot. Btzg II: pl. 2: f. la-c. 1916
  • =Campium foxworthyi Copel in Philip. J. Sc. 37: f. 17, pl. 12. 1928
  • B. foxworthyi Ching in C. Chr., Ind. Fil.: 48. 1934
  • =Campium membranaceum Copel. in Philip. J. Sc. 40: pl. 7. 1928
  • B. membranacea C. Chr., Ind. Fil.: 198. 1934
  • =Campium nigrum Copel. in Philip. J. Sc. 37: f. 14, pl. 10. 1928
  • B. nigra Ching in C. Chr., Ind. Fil.: 49. 1934
  • =Campium pseudoscalpturatum Copel. in Philip. J. Sc. 37: f. 16, pl. 11. 1928
  • B. pseudoscalpturata Ching in C. Chr., Ind. Fil.: 49. 1934
  • =Campium tenuissimum Copel. in Philip. J. Sc. 37: f. 18, pl. 13. 1928
  • B. tenuissima Copel., Fern Fl. Philip.: 259. 1960
  • =Leptochilus simplicifolius Holttum in Gard. Bull. S. S. 4: 409. 1929
  • B. simplicifolia Ching in C. Chr., Ind. Fil.: 50. 1934
  • =B. edanyoi Copel. in Philip. J. Sc. 81: pl. 15. 1952
  • =Edanyoa difformis Copel. in Philip. J. Sc. 81: pl. 17. 1952

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