Combretum paraguariense

Primary tabs

Combretum paraguariense

Content

<<<Flowers>Calyx lobes>Apex>Shape

obtuse to subacute1
1. 004-004-004-001

<<<Flowers>Disk>Margins>Width

ca. 0.3-0.4 mm2
2. 004-007-002-001

the only part of plant densely lepidote3
3. 004-002-002-003

1.7-3.6 mm long4
4. 004-002-002-001

without pedicel-like lower region5
5. 004-002-002-002

<<<Flowers>Stamens>Filaments>Length

ca. 1.5 mm long6
6. 004-006-003-001

<<<Fruits>Pseudostipe>Ridges>Width

0-0.5 cm wide7
7. 005-005-003-001

<<<Fruits>Pseudostipe>Shape>Angle number

4-angled8
8. 005-005-002-001

<<<Fruits>Pseudostipe>Shape>Wing number

4-winged9
9. 005-005-002-002

<<<Leaves>Veins>Secondary veins>Secondary vein pair number

4-7 pairs10
10. 002-006-001-001

<<Flowers>Calyx lobes>Apex

apex ;11
11. 004-004-004

<<Flowers>Calyx lobes>Growth form

erect12
12. 004-004-001

<<Flowers>Calyx lobes>Length

up to 0.6 mm long13
13. 004-004-003

<<Flowers>Calyx lobes>Shape

broadly triangular, very short14
14. 004-004-002

<<Flowers>Disk>Hairs

densely pubescent15
15. 004-007-001

<<Flowers>Disk>Margins

with rather narrow () free margins16
16. 004-007-002

pedicel-like above ovary, , ,17
17. 004-002-002

<<Flowers>Lower hypanthium>Shape

narrow18
18. 004-002-001

<<Flowers>Stamens>Filaments

filaments , each anther with rather inconspicuous caruncle between pollen-sacs on inner side;19
19. 004-006-003

<<Flowers>Stamens>Relative dimensions

exceeding hypanthium by up to 0.6 mm20
20. 004-006-002

<<Flowers>Stamens>Stamen number

421
21. 004-006-001

<<Flowers>Style>Length

1.7-2.3 mm long22
22. 004-008-001

<<Flowers>Upper hypanthium>Dimensions

1.3-2 x 1.6-2.3 mm23
23. 004-003-002

<<Flowers>Upper hypanthium>Hairs

moderately lepidote on outside, usually sparsely pubescent inside24
24. 004-003-003

<<Flowers>Upper hypanthium>Shape

deeply cupuliform25
25. 004-003-001

<<Fruits>Apex>Shape

obtuse to narrowly retuse and often apiculate26
26. 005-003-001

<<Fruits>Base>Shape

obtuse to narrowly rounded27
27. 005-004-001

<<Fruits>Pseudostipe>Length

0-0.05 cm long28
28. 005-005-001

<<Fruits>Pseudostipe>Ridges

stiff ridges29
29. 005-005-003

<<Fruits>Pseudostipe>Shape

to narrowly30
30. 005-005-002

<<Inflorescences>Ultimate branches>Hairs

its axis moderately to densely lepidote31
31. 003-003-002

<<Inflorescences>Ultimate branches>Length

1-12 cm long32
32. 003-003-001

<<Leaves>Apex>Shape

abruptly shortly to moderately acuminate33
33. 002-004-001

<<Leaves>Base>Hairs

very sparsely to moderately lepidote below, extremely sparsely to sparsely lepidote above, especially on midvein34
34. 002-005-002

<<Leaves>Base>Shape

rounded to broadly cuneate35
35. 002-005-001

<<Leaves>Petiole>Hairs

sparsely to moderately lepidote36
36. 002-007-002

<<Leaves>Petiole>Length

0.5-2 cm long37
37. 002-007-001

<<Leaves>Veins>Secondary veins

secondary veins ;38
38. 002-006-001

<<Perianth>Marginal cells>Marginal cell number

25-4039
39. 006-003-001

<<Wood>Liana habit>Length

20 m40
40. 001-001-001

<<Wood>Shrub habit>Height

to 5 m41
41. 001-002-001

<Flowers>Calyx lobes

calyx lobes , , ,42
42. 004-004

<Flowers>Disk

disk , bulging between bases of filaments, , leaving very narrow gap around style-base;43
43. 004-007

<Flowers>Length

3-5.6 mm long44
44. 004-001

<Flowers>Lower hypanthium

lower hypanthium and ,45
45. 004-002

<Flowers>Petals

petals absent;46
46. 004-005

<Flowers>Stamens

stamens , opposite each calyx-lobe, ,47
47. 004-006

<Flowers>Style

style , slightly exserted or not so.48
48. 004-008

<Flowers>Upper hypanthium

upper hypanthium , , ;49
49. 004-003

<Fruits>Apex

apex ,50
50. 005-003

<Fruits>Base

base ,51
51. 005-004

<Fruits>Dimensions

2.2-5 x 0.7-2 cm52
52. 005-001

<Fruits>Pseudostipe

pseudostipe , , .53
53. 005-005

<Fruits>Shape

narrowly oblong to narrowly elliptic-oblong in side view54
54. 005-002

<Inflorescences>Length

4-15 cm long55
55. 003-002

<Inflorescences>Position

in leaf-axils or terminal on lateral branches56
56. 003-001

<Inflorescences>Ultimate branches

ultimate branches and very slender,57
57. 003-003

<Leaves>Apex

apex ,58
58. 002-004

<Leaves>Base

base , ;59
59. 002-005

<Leaves>Dimensions

3.5-14.5(-20) x 1.5-7.5(-9) cm60
60. 002-003

<Leaves>Petiole

petiole ,61
61. 002-007

<Leaves>Shape

elliptic or elliptic-oblong to broadly or narrowly so62
62. 002-002

<Leaves>Texture

chartaceous to subcoriaceous63
63. 002-001

<Leaves>Veins

venation eucamptodromous to brochidodromous,64
64. 002-006

<Perianth>Colour

pale brown65
65. 006-002

<Perianth>Diameter

ca. 75-150 µm diam.66
66. 006-001

<Perianth>Marginal cells

marginal cells .67
67. 006-003

<Wood>Hairs

‘combretaceous hairs’ (only inside flowers) and peltate scales present.68
68. 001-003

<Wood>Liana habit

Woody liana to at least69
69. 001-001

<Wood>Shrub habit

less often shrub70
70. 001-002

Description

Woody liana to at least 20 m, less often shrub to 5 m; ‘combretaceous hairs’ (only inside flowers) and peltate scales present. Leaves opposite or rarely in threes, chartaceous to subcoriaceous, elliptic or elliptic-oblong to broadly or narrowly so, 3.5-14.5(-20) x 1.5-7.5(-9) cm, apex abruptly shortly to moderately acuminate, base rounded to broadly cuneate, very sparsely to moderately lepidote below, extremely sparsely to sparsely lepidote above, especially on midvein; venation eucamptodromous to brochidodromous, secondary veins 4-7 pairs; petiole 0.5-2 cm long, sparsely to moderately lepidoteInflorescence branched or not, in leaf-axils or terminal on lateral branches, 4-15 cm long, ultimate branches 1-12 cm long and very slender, its axis moderately to densely lepidote. Flowers 3-5.6 mm long; lower hypanthium narrow and pedicel-like above ovary, 1.7-3.6 mm long, without pedicel-like lower region, the only part of plant densely lepidote,upper hypanthium deeply cupuliform, 1.3-2 x 1.6-2.3 mm, moderately lepidote on outside, usually sparsely pubescent inside; calyx lobes erect, broadly triangular, very short, up to 0.6 mm long, apex obtuse to subacute; petals absent; stamens 4, opposite each calyx-lobe, exceeding hypanthium by up to 0.6 mm, filaments ca. 1.5 mm long, each anther with rather inconspicuous caruncle between pollen-sacs on inner side; disk densely pubescent, bulging between bases of filaments, with rather narrow (ca. 0.3-0.4 mm) free margins, leaving very narrow gap around style-base; style 1.7-2.3 mm long, slightly exserted or not so. Fruit sparsely to moderately lepidote, 2.2-5 x 0.7-2 cm, narrowly oblong to narrowly elliptic-oblong in side view, apex obtuse to narrowly retuse and often apiculate, base obtuse to narrowly rounded, pseudostipe 0-0.05 cm long, 4-angled to narrowly 4-winged, stiff ridges 0-0.5 cm wide. Scales ca. 75-150 µm diam., pale brown, marginal cells 25-40.

Distribution

French Guiana present, Guianas present, Paraguay and São Lorenzo rivers in Mato Grosso present, S Brazil present, Southern America: Bolivia (Bolivia present); Colombia (Colombia present); Ecuador (Ecuador present); Peru (Peru present); Venezuela (Venezuela present), throughout the Amazon river-basin from its mouth present
Scattered throughout the Amazon river-basin from its mouth, extending to French Guiana, Venezuela, Colombia, Ecuador, Peru and Bolivia, and S to the Paraguay and São Lorenzo rivers in Mato Grosso (S Brazil); 2 collections from the Guianas (FG: 4).

Flowers

Flowers ;71
71. 004

Fruits

Fruit sparsely to moderately lepidote, , ,72
72. 005

Individuals Association

Oyapock R., Camopi R., Oldeman 2637 Oldeman 1791 Oyapock R., Cr. Gabaret, Oldeman B-1531 Oldeman 2652

Inflorescences

Inflorescence branched or not, , , .73
73. 003

Leaves

Leaves opposite or rarely in threes, , , ,74
74. 002

Perianth

Scales , ,75
75. 006

Wood

, ;76
76. 001