Primary tabs



often coloured1
1. 006-001-002


2. 006-001-001


cotyledons thick, equal or unequal.3
3. 006-002-001


at first completely covered with peltate bracts4
4. 003-001-002


(sub)globose, hemispheric, turbinate or convexly discoid5
5. 003-001-001


fully amplexicaul and connate, or semi-amplexicaul and free6
6. 002-001-001

<<Pistillate flowers>Stigma>Shape

filiform to band-shaped7
7. 005-003-002

<<Pistillate flowers>Stigma>Stigma number

8. 005-003-001

<<Staminate flowers>Stamens>Stamen number

9. 004-003-001


with the enlarged, and receptacle10
10. 006-001


seed large, without endosperm,11
11. 006-002


receptacle ,12
12. 003-001


blade entire, pinnately veined.13
13. 002-002


stipules ;14
14. 002-001

<Pistillate flowers>Flower number

one to several15
15. 005-001

<Pistillate flowers>Ovary

ovary adnate to the perianth;16
16. 005-002

<Pistillate flowers>Stigma

stigmas , .17
17. 005-003

<Staminate flowers>Flower number

several to numerous18
18. 004-001

<Staminate flowers>Perianth

perianth well-developed, vestigial or lacking;19
19. 004-002

<Staminate flowers>Pistillode

pistillode absent.20
20. 004-004

<Staminate flowers>Stamens

stamens ;21
21. 004-003


Trees, monoecious or dioecious. Leaves alternate and distichous; stipules fully amplexicaul and connate, or semi-amplexicaul and free; blade entire, pinnately veined. Inflorescences in the leaf axils, bisexual or unisexual, pedunculate, receptacle (sub)globose, hemispheric, turbinate or convexly discoid, at first completely covered with peltate bracts. Staminate flowers several to numerous, on the surface of the receptacle; perianth well-developed, vestigial or lacking; stamens 4-1; pistillode absent. Pistillate flowers one to several, embedded in the receptacle; ovary adnate to the perianth; stigmas 2, filiform to band-shaped. Fruit(s) forming a drupaceous whole with the enlarged, fleshy and often coloured receptacle; seed large, without endosperm, cotyledons thick, equal or unequal.


Guianas present, Neotropics present
Neotropics 13 species; the Guianas 7 species.


Fruit(s) forming a drupaceous whole ;22
22. 006


Trees, monoecious or dioecious.23
23. 001


Inflorescences in the leaf axils, bisexual or unisexual, pedunculate, .24
24. 003


Leaves alternate and distichous;25
25. 002

Pistillate flowers

Pistillate flowers , embedded in the receptacle;26
26. 005

Staminate flowers

Staminate flowers , on the surface of the receptacle;27
27. 004


Vessels diffuse, solitary (20-70%) and in short radial multiples and irregular clusters of 2-4, round to oval, 3-16 per sq. mm, diameter 70-185 μm. Vessel-member length: 400-600 μm. Perforations simple. Inter- vascular pits alternate, round, oval or polygonal, 3-13 μm. Vessel-ray and vessel-parenchyma pits variable, irregular in size and shape, half- bordered. Thin-walled tyloses often present, sometimes thick-walled tyloses occur.
Rays uniseriate and 2-4(7)-seriate, 5-9 per mm, up to 370-610 μm high. Heterogeneous, composed of procumbent cells, except for the uniseriate margins of 1-2 rows of square and upright cells. Rhombic crystals absent to extremely abundant in the marginal ray cells. Radial latex tubes common.
Parenchyma paratracheal, vasicentric-aliform with narrow wings, which are sometimes arc-shaped, occasionally confluent, sometimes banded, mostly unilateral. Strands of 2-5 cells. Rhombic crystals scarce.
Fibres non-septate, lumen 3-16 μm, walls 2-3(5) μm. Pits simple, small, confined to the radial walls. Gelatinous fibres scarce. Length: 980-1600 μm. F/V ratio: 2.0-3.4.

Table 1. Some wood anatomical features of species of Brosimum
VESSELSfrequency (sq. mm)diameter (μm)pit size (μm)tyloses
B. acutifolium4- 9130-1857-13-
B. alicastrum9-1670-1205- 7(t,T)
B. guianense6-1090-1304- 6T
B. lactescens7-16100-1353- 4t
B. parinarioides3-6140-1709-11-
B. rubescens4- 5125-1805- 8T
B. utile31859-13-
Legenda:T = Thick-walled; t = thin-walled
+ = present; — = absent
between brackets = occasionally present
PARENCHYMAconfluent-wavyregular bandsunilat. arrang.arc. sh. wingsradial latex tub.rhombic crystals
B. acutifolium--+-+r(a)
B. alicastrum+++-+rr(a)
B. guianense(+)-+(+)(+)rr(a)
B. lactescens(+)-(+)-+(r)
B. parinarioides-(+)(+)++-
B. rubescens-(+)(+)(+)+r(a)
B. utile(+)(+)(+)-+-
Legenda:+ = present; — = absent
r = in marginal ray cells; rr = abundant in marginal ray cells
a = in axial parenchyma
between brackets = occasionally present

Wood observation species

B. acutifolium subsp. acutifolium, B. acutifolium subsp. obovatum, B. alicastrum subsp. bolivarense, B. guianense, B. lactescens, B. parinarioides subsp. parinarioides, B. rubescens, B. utile subsp. ovatijolium