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<<<Pistillate inflorescences>Seeds>Cotyledons>Shape

thick and equal1
1. 005-006-001-001


2. 002-001-001

<<Pistillate inflorescences>Flowers>Flower number

several to numerous3
3. 005-001-001

<<Pistillate inflorescences>Flowers>Shape

connate and the outer ones adnate to the receptacle4
4. 005-001-002

<<Pistillate inflorescences>Seeds>Cotyledons

cotyledons .5
5. 005-006-001

<<Pistillate inflorescences>Stigma>Shape

filiform to vittiform. lnfructescences hemispherical to globose6
6. 005-004-002

<<Pistillate inflorescences>Stigma>Stigma number

7. 005-004-001

<<Pistillate inflorescences>Tepals>Shape

the free parts conical to cushion-shaped8
8. 005-002-002

<<Pistillate inflorescences>Tepals>Tepal number

9. 005-002-001

<<Staminate inflorescences>Involucral bracts>Shape

often more or less petaloid10
10. 004-002-001

<<Staminate inflorescences>Stamens>Stamen number

11. 004-004-001

<<Staminate inflorescences>Tepals>Tepal number

12. 004-003-001


13. 003-001


blade entire, pinnately veined.14
14. 002-003


distichous on the branches15
15. 002-001


stipules fully amplexicaul, free;16
16. 002-002

<Pistillate inflorescences>Flowers

flowers , ;17
17. 005-001

<Pistillate inflorescences>Fruits

fruits embedded in the pulpy receptacle;18
18. 005-005

<Pistillate inflorescences>Ovary

ovary adnate to the perianth;19
19. 005-003

<Pistillate inflorescences>Seeds

seed large, without endosperm,20
20. 005-006

<Pistillate inflorescences>Stigma

stigmas , , covered with the hardened, free parts of the tepals and the inner involucral bracts;21
21. 005-004

<Pistillate inflorescences>Tepals

tepals , ;22
22. 005-002

<Staminate inflorescences>Inflorescence number

often up to 4 together23
23. 004-001

<Staminate inflorescences>Involucral bracts

the inner involucral bracts , covering the flowers before anthesis24
24. 004-002

<Staminate inflorescences>Pistillode

pistillode absent.25
25. 004-005

<Staminate inflorescences>Stamens

stamens ;26
26. 004-004

<Staminate inflorescences>Tepals

tepals , free;27
27. 004-003


Trees, dioecious. Leaves alternate, distichous on the lateral branches; stipules fully amplexicaul, free; blade entire, pinnately veined. Inflorescences on spurs in the leaf axils, unisexual, discoid-capitate, involucrate. Staminate inflorescences often up to 4 together, pedunculate, the inner involucral bracts often more or less petaloid, covering the flowers before anthesis; tepals 3-7, free; stamens 2-4; pistillode absent. Pistillate inflorescences solitary, (sub)sessile; flowers several to numerous, connate and the outer ones adnate to the receptacle; tepals 3-7, the free parts conical to cushion-shaped; ovary adnate to the perianth; stigmas 2, filiform to vittiform. lnfructescences hemispherical to globose, covered with the hardened, free parts of the tepals and the inner involucral bracts; fruits embedded in the pulpy receptacle; seed large, without endosperm, cotyledons thick and equal.


Guianas present, Neotropical present
Neotropical, 20-25 species, some of them not yet described; in the Guianas 2 (or 3?) species.


Trees, dioecious.28
28. 001


Inflorescences on spurs in the leaf axils, unisexual, , involucrate.29
29. 003


Leaves alternate, ;30
30. 002

Pistillate inflorescences

Pistillate inflorescences solitary, (sub)sessile;31
31. 005

Staminate inflorescences

Staminate inflorescences , pedunculate, ;32
32. 004


Vessels diffuse with a tendency towards a diagonal arrangement, solitary (20-80%) and in short radial multiples of 2-4, angular, round or oval, 20-45 per sq. mm, diameter 60-110 μm. Vessel-member length: 380-475 μm. Perforations simple. Intervascular pits alternate, round, oval to polygonal, 6-9 μm. Vessel-ray and vessel-parenchyma pits larger and irregularly shaped, half-bordered. Thin-walled tyloses occasionally present.
Rays uniseriate and 2-4-seriate, 6-8 per mm, up to 600-1500 μm. Heterogeneous, composed of procumbent cells, except for the uniseriate margins of 1-6 rows of square and/or upright cells. Few sheath cells. Rays sometimes vertically fused. Radial latex tubes present. Rhombic crystals sometimes present in the (marginal) ray cells.
Parenchyma paratracheal, vasicentric-aliform, occasionally locally confluent, the vasicentric ring often incomplete and lacking on the adaxial side. Strands 2-4 cells. Rhombic crystals very scarce.
Fibres septate, lumen 5-10 μm, walls 2-4 μm. Pits simple, small, restricted to the radial walls. Gelatinous fibres occasionally present. Length: 1000-1200 μm. F/V ratio: 2.4-2.8.
Vitreous silica occasionally present in the fibres and parenchyma.

Wood observation species

N. amara, N. glabra, N. guianensis