Besleria laxiflora

<<<Calyx>Corolla>Base>Width

0.3-0.7 cm wide1

1. 005-008-005-001

<<<Calyx>Corolla>Limbs>Width

0.3-0.5 cm wide2

2. 005-008-007-001

<<<Calyx>Corolla>Lobes>Dimensions

ca. 0.2 x 0.1-0.3 cm wide3

3. 005-008-008-003

<<<Calyx>Corolla>Lobes>Growth form

spreading4

4. 005-008-008-001

<<<Calyx>Corolla>Lobes>Shape

ovate5

5. 005-008-008-002

<<<Calyx>Corolla>Margins>Margin type

entire6

6. 005-008-009-001

<<<Calyx>Corolla>Throat>Hairs

outside glabrous, inside with pubescent ring near attachment of filaments and second ring in throat7

7. 005-008-006-002

<<<Calyx>Corolla>Throat>Width

0.4-0.7 cm wide8

8. 005-008-006-001

<<<Calyx>Corolla>Tube>Length

1.2-2 cm long9

9. 005-008-004-002

<<<Calyx>Corolla>Tube>Shape

cylindric10

10. 005-008-004-001

<<<Calyx>Ovary>Style>Hairs

puberulent11

11. 005-010-004-002

<<<Calyx>Ovary>Style>Length

ca. 0.8 cm long12

12. 005-010-004-001

<<<Leaves>Blade>Base>Hairs

above sparsely strigillose on midvein to glabrate, below sparsely strigillose on veins13

13. 003-002-006-001

<<Calyx>Apex>Hairs

outside glabrescent, inside glabrous14

14. 005-007-001

<<Calyx>Corolla>Base

base not spurred or gibbous, , middle slightly ventricose,15

15. 005-008-005

<<Calyx>Corolla>Colour

yellow, orange, salmon red, pink to golden red16

16. 005-008-002

<<Calyx>Corolla>Growth form

erect in calyx17

17. 005-008-001

<<Calyx>Corolla>Length

1.3-2.3 cm long18

18. 005-008-003

<<Calyx>Corolla>Limbs

limb ,19

19. 005-008-007

<<Calyx>Corolla>Lobes

lobes subequal, , , ,20

20. 005-008-008

<<Calyx>Corolla>Margins

margin ;21

21. 005-008-009

<<Calyx>Corolla>Throat

throat slightly constricted, , ,22

22. 005-008-006

<<Calyx>Corolla>Tube

tube , ,23

23. 005-008-004

<<Calyx>Free portion of lobes>Dimensions

0.5-0.7 x 0.2-0.5 cm24

24. 005-005-003

<<Calyx>Free portion of lobes>Growth form

erect25

25. 005-005-001

<<Calyx>Free portion of lobes>Shape

ovate to narrowly lanceolate26

26. 005-005-002

<<Calyx>Lobes>Relative distance

to 1/4 their length27

27. 005-003-001

<<Calyx>Margins>Margin type

entire to minutely fimbriate or ciliate28

28. 005-006-001

<<Calyx>Ovary>Dimensions

0.2-0.4 x 0.2-0.3 cm29

29. 005-010-002

<<Calyx>Ovary>Hairs

glabrous30

30. 005-010-003

<<Calyx>Ovary>Shape

ovoid31

31. 005-010-001

<<Calyx>Ovary>Stigma

stigma stomatomorphic (in the Guianas).32

32. 005-010-005

<<Calyx>Ovary>Style

style , ,33

33. 005-010-004

<<Calyx>Tube>Length

0.6-1.7 cm long34

34. 005-004-001

<<Flowers>Pedicels>Hairs

sparsely strigillose35

35. 004-003-002

<<Flowers>Pedicels>Length

0.8-2.5 cm long36

36. 004-003-001

<<Flowers>Peduncle>Hairs

sparsely strigillose37

37. 004-002-002

<<Flowers>Peduncle>Length

1.5-4.5(-6.5) cm long38

38. 004-002-001

<<Leaves>Blade>Apex

apex acuminate,39

39. 003-002-005

<<Leaves>Blade>Base

base cuneate, .40

40. 003-002-006

<<Leaves>Blade>Dimensions

3-21 x 1-8.5 cm41

41. 003-002-003

<<Leaves>Blade>Margins

margin shallowly toothed,42

42. 003-002-004

<<Leaves>Blade>Shape

elliptic, narrowly oblong, or rarely ovate43

43. 003-002-002

<<Leaves>Blade>Texture

membranous when dry44

44. 003-002-001

<<Leaves>Petiole>Hairs

sparsely strigillose45

45. 003-001-002

<<Leaves>Petiole>Length

1.5-7 cm long46

46. 003-001-001

<Calyx>Apex

apex mucronate, ;47

47. 005-007

<Calyx>Colour

green to yellow or orange48

48. 005-002

<Calyx>Corolla

corolla , , ,49

49. 005-008

<Calyx>Free portion of lobes

free portion of lobes , equal, , ,50

50. 005-005

<Calyx>Lobes

lobes free or rarely connate ,51

51. 005-003

<Calyx>Margins

margin ,52

52. 005-006

<Calyx>Ovary

ovary , , ,53

53. 005-010

<Calyx>Shape

tubular-campanulate54

54. 005-001

<Calyx>Stamens

stamens included, inserted below middle of corolla tube;55

55. 005-009

<Calyx>Tube

tube ,56

56. 005-004

<Flowers>Flower number per inflorescence

1- to several-flowered57

57. 004-001

<Flowers>Pedicels

pedicel , .58

58. 004-003

<Flowers>Peduncle

peduncle , ;59

59. 004-002

<Habit>Height

0.5-4(-7) m tall60

60. 001-001

<Leaves>Blade

blade , , ,61

61. 003-002

<Leaves>Petiole

petiole , ;62

62. 003-001

<Mature berries>Colour

orange or red63

63. 006-001

<Mature berries>Dimensions

0.7-1.2 x 1-1.2 cm64

64. 006-003

<Mature berries>Shape

globose to ovoid65

65. 006-002

<Stems>Growth form

more or less erect66

66. 002-001

<Stems>Hairs

strigose at apex, glabrescent below67

67. 002-002

Calyx

Calyx , ,68

68. 005

Common Name

English (Guyana): turuquaréochuru

Description

Terrestrial suffrutescent herb or low shrub, 0.5-4(-7) m tall. Stem sappy, more or less erect, strigose at apex, glabrescent below. Leaves subequal in a pair; petiole 1.5-7 cm long, sparsely strigillose; blade membranous when dry, elliptic, narrowly oblong, or rarely ovate, 3-21 x 1-8.5 cm, margin shallowly toothed, apex acuminate, base cuneate, above sparsely strigillose on midvein to glabrate, below sparsely strigillose on veins. Flowers umbellate to cymose, in 1- to several-flowered inflorescences; peduncle 1.5-4.5(-6.5) cm long, sparsely strigillose; pedicel 0.8-2.5 cm long, sparsely strigillose. Calyx tubular-campanulate, green to yellow or orange, lobes free or rarely connate to 1/4 their length, tube 0.6-1.7 cm long, free portion of lobes erect, equal, ovate to narrowly lanceolate, 0.5-0.7 x 0.2-0.5 cm, margin entire to minutely fimbriate or ciliate, apex mucronate, outside glabrescent, inside glabrous; corolla erect in calyx, yellow, orange, salmon red, pink to golden red, 1.3-2.3 cm long, tube cylindric, 1.2-2 cm long, base not spurred or gibbous, 0.3-0.7 cm wide, middle slightly ventricose, throat slightly constricted, 0.4-0.7 cm wide, outside glabrous, inside with pubescent ring near attachment of filaments and second ring in throat, limb 0.3-0.5 cm wide, lobes subequal, spreading, ovate, ca. 0.2 x 0.1-0.3 cm wide, margin entire; stamens included, inserted below middle of corolla tube; ovary ovoid, 0.2-0.4 x 0.2-0.3 cm, glabrous, style ca. 0.8 cm long, puberulent, stigma stomatomorphic (in the Guianas). Mature berry orange or red, globose to ovoid, 0.7-1.2 x 1-1.2 cm.

Distribution

Guianas present, Guyana present, Northern America, Southern America: Colombia (Colombia present); Venezuela (Venezuela present), Suriname present

Mexico through Central America to Colombia, Venezuela, Brazil, Guyana and Suriname; > 450 specimens studied, 24 from the Guianas (GU: 13; SU: 11).

Flowers

Flowers umbellate to cymose, in inflorescences;69

69. 004

Habit

Terrestrial suffrutescent herb or low shrub, .70

70. 001

Individuals Association

Suriname, Brokopondo Distr., Brownsberg, Stahel & Gonggrijp 19 Guyana, Upper Takutu-Upper Essequibo Region, NW slopes of Kanuku Mts., Moku-Moku Cr. (Takutu tributary), A.C. Smith 3555 Guyana, Potaro-Siparuni Region, Iwokrama Mts., Clarke 2525 Suriname, Emmaketen, Gongrijp & Stahel 182 Suriname, Tjon-Lim-Sang & van de Wiel 32 Guyana, East Berbice-Corentyne Region, New R., Guppy 370

Leaves

Leaves subequal in a pair;71

71. 003

Mature berries

Mature berry , , .72

72. 006

Phenology

Collected in flower in , and in fruit in .

Stems

Stem sappy, , .73

73. 002

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