2-locular3

4- or 5-lobed5

ovule , basal in single carpels, axile in syncarpous ovaries, campylotropous,7

style, if present, as many as carpels, usually free,9

filaments in , sometimes borne on a hypogynous disk,11

united below in a perianth13

embryo curved.15

1-numerous17

stamens , free or the filaments united at the base,19

seeds ,21

terminal or pseudoaxillary23

stipules present or absent.25

Fruit a berry, achene, capsule, samara or drupe;27

Guyana, Schomburgk, Ro. 470 (K) Granville, J. J. de, et al. 8703 Suriname, Everaarts, A.P. 494 Guyana, Cruz, J.S. de la 2501 Taverne, B. 88 Feuillet, C. 3945 Suriname, Jonker-Verhoef, A.M.E. & F.P. Jonker 207 Suriname, BW 261 Suriname, Everaarts, A.P. 1088 Guyana, Schomburgk, Ri. 47 Bordenave, B. 349 Suriname, Went, F.A.F.C. 230 Suriname, Wullschlägel, H.R. 446 Guyana, Cooper, A. 32 Suriname, Rombouts, H.E. 908 Guyana, Appun, C.F. 327 Mori, S. A. et al. 23818 Guyana, Ramsammy, J.R. 9 Suriname, Samuels, J.A. 115 Granville, J. J. de, et al. 7826 Broadway, W.E. 737 Guyana, Jansen-Jacobs, M.J. et al. 3835 Suriname, Kalpoe, C.D. UVS 16588 Kodjoed, J.F. 92 Suriname, Irwin, H.S. et al. 55120 Suriname, Kramer, K.U. & W.H.A. Hekking 2154 Skog, L. & C. Feuillet 7300 Suriname, Jonker-Verhoef, A.M.E. & F.P. Jonker 29 Moretti, C. 1294 Feuillet, C. 90 Suriname, Tryon, R.M. 5672 Suriname, Wessels Boer, J.G. 568 Suriname, Hall, C.J.J. van 100 Service Forestier s.n. Cremers, G.A. et al. 9759 Guyana, Steege, H. ter et al. 224 Suriname, Smith-Naarendorp, E.S. LBB 12904 Oldeman, R.A.A. B-3892 Guyana, Harrison, S.G. & R. Persaud 1072 Suriname, Sailings U-94 Suriname, Focke, H.C. 285 Suriname, Hekking, W.H.A. 1124 Guyana, Pipoly, J.J. et al. 7924 Guyana, Jenman, G.S. 2192 Suriname, Pulle, A. 459 Suriname, Irwin, H.S. et al. 54750 Suriname, BW 511 Suriname, Daniëls, A.G.H. & F.P. Jonker 976 Fournet, A. 198 Riera, B. 930 Suriname, Stahel, G. & J.W. Gonggrijp 6324 Suriname, Wessels Boer, J.G. 686 Suriname, Boerboom, J.H.A. 12001 Suriname, Samuels, J.A. 283 Guyana, Gleason, H.A. 194 Suriname, Hekking, W.H.A. 1164 Suriname, Donselaar, J. van 2402 Guyana, Cruz, J.S. de la 3221 Suriname, Florschütz, P.A. et al. 339 Guyana, Jenman, G.S. 5277 Suriname, LBB Suriname, Heyde, N.M. et al. 310 Suriname, Irwin, H.S. et al. 57701 Feuillet, C. 2181 Suriname, Jonker-Verhoef, A.M.E. & F.P. Jonker 199 Suriname, Samuels, J.A. s.n. Suriname, Maguire, B. 24307 Suriname, Schulz, J.P. 8747 Guyana, Hitchcock, A.S. 16587 Oldeman, R.A.A. 1259 Suriname, Sterringa, J.T. LBB 12533 Suriname, Heyde, N.M. et al. 124 Sastre, C. & F. Sastre 282 Guyana, Hoffman, B. 2419 Guyana, Kvist, L.P. et al. 259 Guyana, Fairchild, D. s.n. Oldeman, R.A.A. B-438 Mori, S. A. et al. 18246 Guyana, Botanical Garden s.n. Oldeman, R.A.A. B-4019 Prévost, M.F. et al. 746 Bordenave, B. 294 Guyana, Hitchcock, A.S. 17383 Guyana, Fanshawe, D.B. 2450 Guyana, Henkel, T.W. 5988 Guyana, Pipoly, J.J. et al. 9716 Suriname, Heyde & Lindeman 124 Guyana, Cruz, J.S. de la 2132 Prévost, M.F. et al. 717 Guyana, Hitchcock, A.S. 17115 Suriname, Reijenga, Th.W. 975 Guyana, Gillespie, L.J. et al. 1027 Guyana, Smith, A.C. 3650 Guyana, Schomburgk, Ri. 203 Guyana, Davis, D.H. 893 Suriname, Stoffers, A.L. 18B Guyana, Appun, C.F. 2459 Alexandre, D.Y. 164 Guyana, Henkel, T.W. 630 Guyana, Kvist, L.P. et al. 5 Guyana, Stoffers, A.L. & A.R.A. Görts-van Rijn et al. 193 Broadway, W.E. 222 Oldeman, R.A.A. 3120 Suriname, Hugh-Jones, D.Ll. 25 Guyana, Sandwith, N.Y. 1566 Alexandre, D.Y. 486 Guyana, Cruz, J.S. de la 3650 Guyana, Hitchcock, A.S. 16608 Suriname, Heyde 626 Guyana, Jenman, G.S. 6524 Suriname, Boerboom, J.H.A. LBB 9179 Guyana, Schomburgk, Ro. ser. I, 229 Guyana, Jenman, G.S. 503 Guyana, Clarke, H.D. 978 Billiet, F.P. & B. Jadin 1165 Suriname, BW 521 Mori, S. A. et al. 18639 Guyana, Grewal, M.S. & R. Persaud 94 Guyana, Schomburgk, Ro. ser. I, 116 Suriname, Rombouts, H.E. 625 Suriname, Stockdale, F.A. s.n. Guyana, Cruz, J.S. de la 4601 Alexandre, D.Y. 392 Larpin, D. 453 Guyana, Cruz, J.S. de la 4033 Guyana, Schomburgk, Ro. ser. I, 835 Guyana, Gleason, H.A. 696 Guyana, Irwin, H.S. 788 Jacquemin, H. 1498 Suriname, Daniëls, A.G.H. & F.P. Jonker 1226 Guyana, Cruz, J.S. de la 2085 Wittingthon, V. 58 Guyana, Hahn, W. et al. 3903 Guyana, Gillespie, L.J. et al. 1313 Poiteau, P.A. s.n. Alexandre, D.Y. 338 Guyana, FD 5186 Guyana, Irwin, H.S. 237 Fleury, M. 492 Feuillet, C. 663 Prévost, M.F. et al. 892 Guyana, Jenman, G.S. 5151 Guyana, Pipoly, J.J. et al. 8307 Moretti, C. 734 Fournet, A. 139 Kodjoed, J.F. 99 Suriname, Lindeman, J.C. 1105 Alexandre, D.Y. 68 Garnier, F.A. 102 without coll. 1865 Fleury, M. 451 Suriname, Kramer, K.U. 115 Grenand, P. 53 Granville, J. J. de, et al. 2532 Guyana, Hahn, W. et al. 5676 Oldeman, R.A.A. 1651 Grenand, P. 201 Guyana, Cruz, J.S. de la 3639 Guyana, Lee, G. & R. Persaud 18 Suriname, Heyde, N.M. et al. 626 Suriname, Soeprato B11 Suriname, Doesburg, P.H. van 109 Leclerc, A. 0051 Guyana, Jansen-Jacobs, M.J. et al. 1546 Suriname, UVS Suriname, Versteeg, G.M. 500 Guyana, Schomburgk, Ro. ser. I, 661 Feuillet, C. 2502 Grenand, P. 1670 Moretti, C. 585 Guyana, Jansen-Jacobs, M.J. et al. 4250 Guyana, Schomburgk, Ro. s.n. Guyana, Roberston, K.R. & D.F. Austin 282 Guyana, Jenman, G.S. 7849 Guyana, Gleason, H.A. 55 Suriname, Smith-Naarendorp, E.S. 12915 Moretti, C. 119 Guyana, Henkel, T.W. 4150 Suriname, Stockdale, F.A. s.n. Guyana, Gleason, H.A. 826 Guyana, Jansen-Jacobs, M.J. et al. 1005 Hoff, M. 6252 Suriname, Focke, H.C. 223 Guyana, Jenman, G.S. 2013 Suriname, Wullschlägel, H.R. 1683 Suriname, Samuels, J.A. 65 Guyana, Mell, C.D. & R.C. Mell 234 Suriname, Pons, T.L. LBB 12638 Sauvain, M. 768 Guyana, Cruz, J.S. de la 2480 Guyana, Warren, L.B. s.n. Suriname, Sterringa, J.T. LBB 12424 Suriname, Tresling, J.H.A.T. 147 Guyana, Cruz, J.S. de la 3130 Guyana, Mutchnick, P. 1501 Guyana, Schomburgk, Ro. s.n. Guyana, Tate, G.H.H. 299 Suriname, Florschütz, P.A. et al. 989 Suriname, Samuels, J.A. 8 (NY) Suriname, Versteeg, G.M. 618 Guyana, Archer, W.A. 2256 Guyana, Steege, H. ter et al. 439 Prévost, M.F. et al. 1356 Weitzman, A. & W. Hahn 318 Guyana, Jansen-Jacobs, M.J. et al. 236 Guyana, Schomburgk, Ro. s.n. Guyana, Cruz, J.S. de la 3548 Guyana, Stockdale, F.A. s.n. Suriname, Samuels, J.A. 8 (K) Guyana, Jansen-Jacobs, M.J. et al. 4733 Granville, J. J. de, et al. 1150 Granville, J. J. de, et al. 3649 Guyana, Cruz, J.S. de la 4481 Guyana, Archer, W.A. 2295 Alexandre, D.Y. 294 Guyana, Omawale & R.N. Persaud 217 Suriname, Samuels, J.A. s.n. Mori, S. A. et al. 21186 Deward, G. 173 Hoff, M. 6083 Guyana, Irwin, H.S. 797 Guyana, McDowell, T. et al. 2168 Suriname, Focke, H.C. 885 Guyana, Cruz, J.S. de la 2093 Guyana, Schomburgk, Ro. ser. I, 860 Granville, J. J. de, et al. 10722 Oldeman, R.A.A. 3151 Feuillet, C. 1334 Lescure, J.P. 310 Suriname, Stahel, G. & J.W. Gonggrijp BW 6308 Guyana, Cruz, J.S. de la 1143 Suriname, Lindeman, J.C. & A.L. Stoffers et al. 7 Suriname, Lanjouw, J. 395 Jacquemin, H. 1467 Guyana, Gorinsky, C. 21 Cremers, G.A. et al. 7957 Guyana, McDowell, T. et al. 2627 Burgot, S-J. 4 Feuillet, C. 325 Suriname, Wessels Boer, J.G. 1175 Guyana, Roberston, K.R. & D.F. Austin 291 Suriname, Samuels, J.A. s.n. Suriname, Lanjouw, J. & J.C. Lindeman 2331 Grenand, P. 2875 Mori, S. A. et al. 19065 Suriname, Lanjouw, J. & J.C. Lindeman 1814 Sastre, C. & F. Sastre 4041 Suriname, Boerboom, J.H.A. 9196 Jacquemin, H. 2842 Suriname, Versteeg, G.M. 881 Gely, A. 43 Mori, S. A. et al. 23295 Service Forestier 6028 Guyana, Gleason, H.A. 18 Guyana, Appun, C.F. 1787 Grenand, P. 1117 Chérubini, B. 20

Leaves alternate, simple, entire, usually petiolate;30

Included phloem present in tangential concentric zones, consisting of bundles of xylem and phloem, separated by conjunctive parenchyma (); bands sometimes anastomosing, 200-400 μm wide, 5-6 per cm. Many cells filled with rhombic crystals or long styloids.
Vessels round to oval, solitary and in small multiples, diffuse, regularly distributed or lacking in narrow tangential zones at the outer side of the phloem bands; diameter 40-125 μm, small and large vessels intermingled; 25-40 vessels per mm². Perforations simple. Intervascular pits oval, 4 x 6 μm. Vessel-ray pits similar to intervessel pitting. Vessel member length 300-400 μm.
Rays 1-5-seriate, the multiseriates with short to long uniseriate margins; multiseriate parts up to 750 (2000) μm high, often slightly irregularly built, mainly composed of procumbent cells, the larger rays with sheath cells; margins like uniseriate rays composed of procumbent, square and upright cells; up to 1250 μm, 8 cells high, 5-8 per mm.
Parenchyma very scanty paratracheal as fusiform cells or 2-celled strands; strands 120-150 μm high; as conjunctive tissue in the alternating with the islands of included phloem.
Fibres with moderately thick-walled, walls 3-4 μm, lumina up to 15 μm wide. Pits simple, slit-like, numerous on radial walls, scanty on tangential walls.
Crystals of different size and shape, varying from rhombic and fusiform crystals to styloids in the included phloem (Carlquist 1988).
In the Guianas, three genera occur that are known to include woody species. The only species represented in the Utrecht wood collection is the scandent shrub Seguieria americana.
As far as known the wood is of no commercial value.