Piper humistratum


Creeping herb or subshrub to 0.35 m tall. Stem villous often with white hairs. Petiole 1-2(-5.5) cm long, villous; blade discolourous, not scabrous, not conspicuously glandular-dotted, elliptic to broadly elliptic, 9-15(-18) x 5-7(-8.5) cm, apex obtuse or acutish, base unequally attached to petiole difference 0.2-0.5 cm, obliquely rounded, glabrous above, veins brown-villous below; pinnately veined, secondary veins 3-5 per side, originating from lower 3/4 or more of primary vein, anastomosing, impressed above, prominent below like tertiary veins, those loosely reticulate. Inflorescence erect; peduncle 1.5-2.5 cm long, villous; spike 2-4 cm long, green or yellow or "rosish", not apiculate; floral bracts glabrous, slightly cucullate. Infructescence occasionally pendent; fruits oblongoid, sunken, glabrous, stigmas 4, sessile.


Guianas: present Southern America:, Brazil North (Amapá present) Terr. Roraima: present
The Guianas and Brazil (Terr. Roraima, Amapá); 51 collections studied (GU: 1; SU: 12; FG: 38).

Common Name

English (French Guiana): zaõ saapatu


I agree with Steyermark and Howard (in ) that Quebitea guianensis, nicely depicted in Aublet, belongs to Piper humistratum.


Piper humistratum Görts & K.U. Kramer in Proc. Kon. Ned. Akad. Wetensch. C: t. 1. 1966 sec. C.A. Todzia & al. 2006
    • Holotype: Between Coppename R. and Emma Range, Wessels Boer 1469.
  • =Quebitea guianensis Aubl. in Pl. Hist. Guiane 2: Table Noms Latins 25, 4: t. 327. 1775
    • Holotype: Aublet s.n.



CountryDateCollector + collecting numberHerbariaTypeScanDerivatives
Aublet s.n.
Citation: Aublet s.n.

Specimen summary: Aublet s.n.
Holotype of Quebitea guianensis Aubl.

Wessels Boer 1469
Citation: Between Coppename R. and Emma Range, Wessels Boer 1469.

Specimen summary: Between Coppename R. and Emma Range, Wessels Boer 1469.
Holotype of Piper humistratum Görts & K.U. Kramer

de Granville et al. 11026
Citation: de Granville et al. 11026.

Guyana, Republic ofvan Andel 2430
Citation: Guyana, Republic of, NW Distr., Morucca R., Kabura Backdam, van Andel 2430.

Mori et al. 14909
Citation: Mori et al. 14909.

Freiberg 216
Citation: Saül area, Freiberg 216.

Suriname, Republic ofPlotkin 610
Citation: Suriname, Republic of, SE Suriname near Tapoc, Plotkin 610.

Suriname, Republic ofAcevedo 5939
Citation: Suriname, Republic of, Tumac Humac Mts., Mt. Telouakem, Acevedo 5939.