Piper adenandrum


Shrub or treelet, 1.5 m tall, pubescent or villous, hairs to 0.5 mm long; upper internodes densely glandular. Petiole 0.2-0.5(-1) cm long, vaginate to middle; blade not conspicuously glandular-dotted, lance-elliptic or elliptic-ovate, 8-16.5 x 3-6(-7) cm, margin ciliate, apex long-acuminate, base unequally attached to petiole difference 0-0.2 cm, acute or obtusish, glabrous or pubescent above, (densely) crisp-pubescent or villous beneath; pinnately veined, secondary veins 5-6 per side, originating from lower 3/4 of primary vein. Inflorescence pendent; peduncle 0.5-1.1 cm long, sometimes reddish tinged, crisp-pubescent or villous; spike 1-5 cm long, green to pale or dark yellow, apiculate; rachis glabrous; floral bracts densely marginally fringed; anthers laterally dehiscent. Fruits separate at maturity, depressed globose, 2.5-3(-4) mm in diam., verruculose; stigmas 3, sessile or fruits somewhat stylose.


Guianas: present Southern America:, Venezuela (Venezuela present)
Venezuela and the Guianas; 81 collections studied (GU: 14; SU: 5; FG: 62).

Common Name

Boni (French Guiana): ampukuwiwii; English (Guyana, Republic of): tona tonakeng; English (Suriname, Republic of): katun uwi


Piper adenandrum and P. avellanum had been considered conspecific by Kramer & Görts-van Rijn (1968: 418). There are, however, several differentiating characters: P. adenandrum has long-acuminate blades, indument of spreading, to 0.5 mm long hairs and distinctly separate fruits, 2.5-3(-4) mm in diam., whereas P. avellanum blades are (short) acuminate or acute, the hairs are crisp, 0.2 mm long, and the fruits 2 mm in diam., not distinctly separate.


Piper adenandrum (Miq.) C. DC. in A. DC., Prodr. 16: 273. 1869 sec. C.A. Todzia & al. 2006
  • Artanthe adenandra Miq., Syst. Piperac.: 515. 1844
    • Holotype: Leprieur 152.
    • Isotype: Leprieur 152.
  • =Piper oblongifolium



CountryDateCollector + collecting numberHerbariaTypeScanDerivatives
Guyana, Republic ofClarke 2947
Citation: Guyana, Republic of, Acarai Mts., Clarke 2947.

Guyana, Republic ofClarke 7059
Citation: Guyana, Republic of, Upper Takutu — Upper Essequibo region, Gunn’s, Clarke 7059.

Feuillet 2240
Citation: Kaw Mts., Feuillet 2240.

Leprieur 152
Citation: Leprieur 152.

Specimen summary: Leprieur 152.
Isotype of Artanthe adenandra Miq. Piper adenandrum (Miq.) C. DC.
Holotype of Artanthe adenandra Miq. Piper adenandrum (Miq.) C. DC.

de Granville et al. 8398
Citation: Saül area, foot Mt. Galbao, de Granville et al. 8398.

Suriname, Republic ofSauvain 448
Citation: Suriname, Republic of, Gran Santi, Sauvain 448.

Suriname, Republic ofLindeman & Stoffers et al. 834
Citation: Suriname, Republic of, Lely Mts., Lindeman & Stoffers et al. 834.