1Pinnules of at least the larger pinnae catadromous. Rhizome scales acicular or nearly acicular
1'Pinnules of at least the larger pinnae anadromous. Rhizome scales acicular or not.
2Scales basifixed along broad base, roots borne on all sides of rhizome, sori terminal at the vein endings
2'Scales peltate or pseudopeltate, roots restricted to the ventral side of lateral buds, sori facing midveins at the forking point at the bending of veins, extra-axillary buds lateral to the phyllopodia, or lower lateral and slightly anterior.