| 1 | Pinnules of at least the larger pinnae catadromous. Rhizome scales acicular or nearly acicular |
| 1' | Pinnules of at least the larger pinnae anadromous. Rhizome scales acicular or not. |
| 2 | Scales basifixed along broad base, roots borne on all sides of rhizome, sori terminal at the vein endings |
| 2' | Scales peltate or pseudopeltate, roots restricted to the ventral side of lateral buds, sori facing midveins at the forking point at the bending of veins, extra-axillary buds lateral to the phyllopodia, or lower lateral and slightly anterior. |
