(7b) REGIONAL KEY TO THE SPECIES — MOLUCCAS (20 species)

1Lower leaf surface with persistent dense indumentum; whenever the hairs are short they may be densely interwoven and thus the leaf may seem glabrous.
1'Lower leaf surface either glabrous, glabrescent, with sparse minute scattered hairs, or with a very minute cobweb-like covering (only visible with a lens).
2Twigs l-2.5(-4) mm diam. Leaves 4.5-15 by 1.5-6 cm. Fruits 2.5-3 cm long, with scurfy hairs 0.1-0.2 mm. — Am Is.
2'Twigs 3-6 mm diam. Leaves 20 cm long or more. Fruits either with distinct hairs, 0.5 -1 mm, or hairs scurfy.
3Fruits 4-6.5(-7) cm long, with conspicuous rusty hairs 0.5-1 mm. [Two taxa, with a different distribution, and distinguishable mainly by characters of the male flowers.].
3'Fruits 5-8.5 cm long, hairs dull cinnamon, scurfy, 0.1-0.2 mm. — N Moluccas.
4Male buds 4.5-6 mm long; bracteole mostly caducous.
4'Male buds 3-4.5 mm long; bracteole persistent. — Am Is.
5Twigs distinctly ridged or winged; myrmecophilous. [Fruits subsessile, more or less ovoid, 2 cm long, apiculate, minutely tomentulose.] —Kai Is.
5'Twigs neither ridged nor winged; not myrmecophilous.
6Plants conspicuously glabrous. Flowers not known. — Seram.
6'Indumentum present on terminal leaf bud, twig, and flowers.
7Leaves 6-17(-25) cm long, lower surface with scattered dots and dashes. — Am Is.
7'Leaves of various sizes, without dots and dashes.
8Female buds much narrowed towards the acute apex, the apical part sharp-angular in cross section. Leaves to 30 cm long.
8'Apex of female buds narrowed or not, in cross section circular or at most bluntly angular, but not sharply angular. Leaves small or large, to 40 cm long.
9Buds 5-6 mm long. Leaves below glabrous or with minute scattered hairs. Fruits 3-3.5 cm long, with scattered hairs 0.3-0.5 mm, or glabrescent. — N Moluccas (Sula Is., Bacan, Bum).
9'Buds 10 mm long. Leaves below with dense or scattered scale-like hairs 0.1-0.2 mm. Fruits 5-6 cm long, densely dark brown short-pubescent, hairs 0.2-0.3 mm. — N Moluccas.
10Twigs 3-5(-10) mm diam. Leaves 25-45 cm long.
10'Twigs 1-5 mm diam. Leaves mostly less than 30 cm long (up to 43 cm long in M. nivea). [Fruits 2-5.5 cm long, pubescent, or fruits larger but then glabrous or early glabresent.].
11Twigs 4-7(-10) mm diam.; bark of twigs conspicuously longitudinally cracking, ± flaking. Lower leaf surface papillose (lens!). [Fruits large, (4-)5-8 cm long, with persistent scurfy hairs 0.1-0.2 mm.] — N Moluccas (Bacan).
11'Twigs 5-10 mm diam.; bark of twigs striate, at most finely cracking and finely flaking. Lower leaf surface not papillose. [Female flowers and fruits not known.]— N Moluccas (Bacan).
12Fruits glabrous or largely early glabrescent, hairs 0.1 mm or less. Inflorescences delicate, few-flowered, without or with a slender peduncle. [Leaves elliptic, with few lateral nerves per side.].
12'Fruits with persistent indumentum, hairs may be very short and inconspicuous. Inflorescences with or without a peduncle, with the flowers in a woody scar-covered short-shoot.
13Twigs 2-3.5 mm diam. Leaves ± coriaceous, 9-22 cm long, below late glabrescent. Fruits 4.5-8 cm long. — N Moluccas.
13'Twigs 1-2 mm diam. Leaves chartaceous, 6-13 cm long, below early glabrescent. Fruits 4-6 cm long. — Cultivated (origin Banda).
14Leaves pale below, with scattered dark-coloured small dots, representing hair scars (lens!). — Known in the Moluccas only from a sterile collection from Talaud Is.
14'Leaves not conspicuously pale below, without scattered dots.
15Fruits 3-4 cm long, with conspicuous coarse hairs, (0.5-)l mm.
15'Fruits variable in size, hairs mealy, scurfy, or woolly, shorter, 0.1-0.5(-0.8) mm long.
16Lower leaf surface greyish or whitish, glabrescent, indumentum weak, its stoutest hairs leaving numerous regularly spaced, small, dark dot-like scars (lens!). Fruits short-ellipsoid, 3-4.2 cm long, hairs 0.1-0.5 mm; pericarp thick and woody, 5 mm thick. — N & C Moluccas.
16'Upper and lower leaf surface more concolorous, or, whenever greyish or whitish, without small dots.
17Leaves elliptic to oblong-lanceolate, 14-35 cm long. Fruits ellipsoid, (2.5-)3-3.5 (-4.5) cm long, hairs 0.1-0.2 mm.
17'Leaves generally smaller, elliptic to lanceolate, to 20 cm long. Fruits variable 18 18a. Lower leaf surface not papillose (lens!). Fruits subglobose, 2.5-3 cm long; pericarp 4-8 mm thick, hairs 0.1 mm or less. — N Moluccas.
17''Lower leaf surface papillose. Fruits ellipsoid; pericarp 1-5 mm thick, hairs short or long, 0.1-1 mm. [Four taxa, difficult to segregate without male flowers.].
18Male inflorescences (short) pedunculate; male buds 2.5 mm long. [Fruits ovoid-ellipsoid, 3-4 cm long, with scurfy hairs 0.2 mm.] — N Moluccas (Sula Is., Halmahera, Obi).
18'Male inflorescences (sub)sessile; male buds 3-6 mm long.
19Leaves membranous. [Venation on lower leaf surface distinct. Fruits 2-3.5 cm long, hairs 0.1(-0.2) mm.] Male buds small, 3 mm long — NE Moluccas.
19'Leaves chartaceous. Male buds 4-6 mm long.
20Venation on lower leaf surface generally indistinct; lateral nerves 8-20 per side, rather dense; blade 4-17 cm long. Fruits 1.3—2.6(—3) cm long, hairs 0.1 mm.
20'Venation distinct or not; lateral nerves more lax, 6-12 per side; blade 7-24 cm long. Fruits 2.5-3.5(-3.8) cm long, hairs 0.3-1 mm. — SE Moluccas (Tanimbar Is).