(6) REGIONAL KEY TO THE SPECIES — PHILIPPINES (20 species)

1Lower leaf surface with a persistent dense indumentum; whenever the hairs are small they are densely interwoven.
1'Lower leaf surface either with ± spaced silky hairs, with a very minute cobweb-like covering (to be seen with a lens only), glabrous, or glabrescent.
2Hairs of leaf bud (0.1-)0.5-l mm. [Inflorescences mostly pedunculate. Fruits 2.5-4 cm long, with conspicuous shaggy hairs 0.5-1 (-2) mm.]
2'Hairs of leaf bud shorter.
3Twigs (3-)4-8 mm diam. Leaves (15-)20 cm long or more. Inflorescences a (sub)-sessile scar-covered wart-like brachyblast. Fruits with minute or longish hairs, to 1 mm.
3'Twigs 1-2.5(-4) mm diam. Leaves (5-)8-22 cm long. Inflorescences pedunculate. Fruits with short scurfy hairs, 0.1-0.2 mm.
4Leaves coriaceous, oblong-lanceolate, (15-)20 cm long. [Petiole proportionally long, 20-35 mm.] — NE Luzon: Palanan area.
4'Leaves herbaceous or chartaceous, elliptic-oblong, 15-50 cm long.
5Twigs 5-8 mm diam. Leaves tapered (narrowed) in the lower half, nerves 30-35 per side; lower leaf surface with whitish dense felty hairs 0.1 mm or less, thus seemingly glabrous (lens!). Fruits (5-)6-6.5 cm long.
5'Twigs 3-4(-6) mm diam. Lower leaf surface with yellowish brown hairs 0.1-0.2(-0.5) mm. Fruits 3.5-6.5(-7) cm long.
6Lower leaf surface with minute persistent hairs. Midrib below brown or yellowish. Fruits 5-9 cm long. — Probably Mindanao.
6'Lower leaf surface late glabrescent. Midrib below purplish, contrasting with greyish surface. Fruits 3-7 cm long. — Palawan, Mindoro.
7Buds in apical part sharp-angular. Inflorescences mostly pedunculate and branched. Fruits rather small, 1.5-3 cm, with scattered hairs or glabrescent.
7'Apical part of buds not or not sharply angular. Inflorescences sessile or pedunculate (and branched). Fruits variable.
8Leaves chartaceous or coriaceous, 10-24(-30) cm long; petiole proportionally long, 20-35 mm. Inflorescences (infructescences) sessile. Fruits large, (5-)6-7 cm long [with minute dark rusty scurfy indumentum].
8'Leaves coriaceous, chartaceous, or membranous; petiole proportionally shorter. Inflorescences sessile or pedunculate. Fruits large, or smaller.
9Leaves coriaceous, ll-24(-30) by 3-6 cm; lateral nerves (15-)20 per side. Twigs 4-6 mm diam. — NE Luzon: Palanan area.
9'Leaves chartaceous, 10-15 by 2.5-6 cm; nerves c. 10 per side. Twigs 2-3 mm diam. Petiole proportionally long — Luzon, Biliran.
10Twigs 3-5(-10) mm diam. Leaves large, to 45 cm long. Inflorescences or infructescences with a peduncle. Fruits large, (4-)5-8 cm long, with persistent indumentum.
10'Twigs 1-5 mm diam. Leaves generally smaller, to 20(-30) cm long, and inflorescences pedunculate or (sub)sessile brachyblasts. Whenever fruits are large, they are glabrescent.
11Leaf bud with hairs l(-2) mm. Fruits with hairs 0.3-0.5 mm.
11'Leaf bud with hairs 0.1-0.2(-0.3) mm. Fruits with scurfy hairs 0.1 mm. [Female flowers not known.].
12Twigs 1-2 mm diam. Fruits 3.5-4 cm long, indumentum minute, soon glabrescent. Lower leaf surface distinctly papillose. Bracteole in female flowers well below the perianth. — Palawan.
12'Twigs 2-5 mm diam. Fruits pubescent or glabrescent. Lower leaf surface papillose or not. Bracteole at apex of pedicel.
13Leaves lanceolate, lateral nerves 20-30 per side, flat or slightly raised below (not to be felt with the finger); very distinctly papillose (lens!). Pericarp 10 mm thick.
13'Leaves elliptic or lanceolate, lateral nerves c. 20 per side or fewer, generally raised below; papillae absent or present, conspicuous or not (lens!). Pericarp usually less than 10 mm thick.
14Nerves c. 45° or less to the midrib. Inflorescences (best to be seen in male specimens) pedunculate, the flowers in subumbels.
14'Nerves c. 45° or more to the midrib. Inflorescences with few flowers at the end of a sessile scar-covered, wart-like brachyblast lasting several flowering seasons
15Nerves (45-)60° to the midrib. Lower leaf surface not papillose. — Mindanao: Surigao (Wenzel 3537).
15'Nerves less than 60° to the midrib. Lower leaf surface ± papillose.
16Fruits 3-4.7 cm long, with dull grey-brown scurfy indumentum. Midrib on lower leaf surface (yellow-)brown when dry, not much contrasting
16'Fruits 3-7 cm long, with bright dark rusty or orange-brown scurfy indumentum. Midrib on lower leaf surface drying dark purplish or reddish, much contrasting.
17Leaves below whitish, contrasting with upper surface; lower leaf surface minutely dotted by the scars of fallen larger hairs.
17'The two leaf surfaces ± concolorous or not, not dotted.
18Twigs 1.5-2.5 mm diam. Leaves 20 cm long or less. Fruits ellipsoid, 3.5 cm long, with conspicuous fine-woolly hairs 0.5(-l) mm. Hairs of leaf bud 1 mm. — Mindanao, Samar.
18'Twigs 2-4 mm diam. Leaves 20-40 cm long, greyish below. Fruits ellipsoid, 3.5-4 cm long, hairs 0.1 mm or less. Hairs of leaf bud 0.1 mm. — Mindanao.
19Bark of twigs soon cracking and/or flaking. Fruits 6 cm long, glabrous (glabrescent). [Male flowers not known.] — Basilan.
19'Bark not or only slightly cracking and flaking. Fruits shorter with minute indumentum.
20Twigs 3-4 mm diam., lower down densely set with lenticels. Leaves coriaceous. [Fruits subglobose, 3-4 cm long, hairs 0.1-0.2 mm.] — Mainly a coastal species; 5 Taiwan, Philippines (N Luzon).
20'Twigs (1-)1.5-3 mm diam.; lenticels present or not. Leaves membranous or chartaceous.
21Lenticels of twigs few and inconspicuous. Fruits subglobose or ellipsoid, 3.5-5 cm long, hairs 0.1 mm. Leaves 8.5-23(-28) cm long.
21'Lenticels present, distinct or not. Fruits ellipsoid, (2.5-)3-3.5(-4.5) cm long, hairs 0.1-0.2 mm. Leaves 14-35 cm long. — Mindanao.