(3) REGIONAL KEY TO THE SPECIES — BORNEO (20 species)

1Leaf bud with minute, appressed hairs, 0.1-0.2(-0.3) mm or less.
1'Hairs more than 0.2 mm.
2Lower leaf surface with (sub)persistent dense indumentum (hairs may be very small), old leaves sometimes glabrescent. Fruits with woolly scurfy hairs 0.1 mm. [Inflorescences with distinct peduncle (often short in female).].
2'Lower leaf surface glabrous or early glabrescent or with scattered (not dense) minute hairs. Fruits with variable indumentum, glabrous, or glabrescent.
3Buds (best to be seen in male flowers) ± angular in the upper part. Twigs 1.5-2.5 (-4) mm diam. Leaves (8-)10-22 cm long, lateral nerves 10-17 per side. [Fruits 5-9 cm long, often subsessile. Check also M. smythiesii.].
3'Buds not angular. Twigs 2-8 mm diam. Leaves 15-40 cm long, with 16-33 lateral nerves per side.
4Twigs (4-)5-8 mm diam. Leaves 16-40 cm long, drying dark brown; lateral nerves (20-)23-33 per side. Fruits 6-9 cm long.
4'Twigs 2-3 mm diam. Leaves 15-30 cm long, drying olivaceous; nerves 16-23 per side. Fruits 5-7 cm long.
5Inflorescences pedunculate; perianth largely glabrous (except in M. gigantea and M. papyracea). Fruits glabrous, glabrescent, or with minute scattered hairs; in glabrescent fruits sometimes a minute indumentum remaining in sheltered places near the insertion of the pedicel.
5'Inflorescences a sessile brachyblast, or pedunculate; perianth pubescent. Fruits with persistent dense-scurfy indumentum, or partly glabrescent but with dense indumentum remaining in sheltered places.
6Buds (especially in male flowers) sharply angular in the apical part. Twigs and leaves drying yellowish; lateral nerves 8—13(—15) per side. [Fruits drying dark brown or blackish, often with a green-yellowish exudate, 4.5-7.5 cm long, glabrescent, hairs minute, scattered, 0.1 mm.].
6'Buds rounded. Twigs drying (yellowish) brown, leaves olivaceous or brown; lateral nerves 10-30 per side.
7Lines of interarching of lateral nerves on lower leaf surface (of mature leaves) indistinct.
7'Lines of interarching raised and distinct.
8Bracteole (in male flowers) subpersistent or late caducous. Leaf apex acute-acuminate; lower leaf surface not papillose. Fruits 3-6(-8) cm long, pericarp 3-10 mm thick.
8'Bracteole caducous. Apex of leaves of fertile twigs acute or blunt; lower leaf surface papillose. Fruits 5-8 cm long, pericarp 15-20 mm thick.
9Leaves membranous, chartaceous, or subcoriaceous, 7-24 cm long. Peduncle in female to 1 cm long, 1-2 mm thick. Fruits 4-6(-8) cm long, pericarp 4-10 mm thick, late or early glabrescent. Male buds 4-6(-7) mm long.
9'Leaves thinly membranous, 7-14 cm long. Peduncle in female 1-2.5 cm long, 1.5 mm thick. Fruits 3-4 cm long, early glabrescent; pericarp 3-4 mm thick. Male buds 3 mm long.
10Twigs 2-4 mm diam. Leaves about 3 times longer than broad, 10-30 by 4-9.5 cm, apex acute-acuminate, base rounded to acute; lateral nerves 14-20 per side; lower leaf surface papillose or not. Male buds 3-3.5 mm long. Female buds 2-2.5 mm long. Fruits 4-6 cm long.
10'Twigs 4-8 mm diam. Leaves large and comparatively broad, 16-44 by 8-14.5 (-18) cm, apex blunt or acute, not acute-acuminate, base ± rounded or subcordate; nerves (16-)20-28 per side; lower leaf surface papillose. Male buds 6-8 mm long. Female buds 6 mm long. Fruits 7-9 cm long.
11Inflorescences (and infructescences) with a peduncle, and ± branched.
11'Inflorescences sessile, a short, scar-covered sessile brachyblast. [Leaves chartaceous or thinly coriaceous.].
12Leaves 16-40 cm long; lateral nerves 16-30 per side. Inflorescences large, widely paniculate, 6-18 cm long in male, (1.5—)5—9 cm in female. Fruits 6-8 cm long
12'Leaves 10-30 cm long; lateral nerves 13-20(-22) per side. Inflorescences 2-6 cm long in male, 1.5-2.5 cm long in female. Fruits (4.5-)5-7.5 cm long. [Male buds 3-4 mm long.].
13Twigs (4-)5-8 mm diam. Leaves: lateral nerves (20-)23-30 per side; lines of interarching ± distinct; papillae usually apparent. Male buds 5-7 mm long.
13'Twigs 2.5-5 mm diam. Leaves: lateral nerves 16-24 per side; lines of interarching ± indistinct; papillae not apparent. Male buds 4(-4.5) mm long
14Twigs 5-10 mm diam. [Bark usually conspicuously cracking and flaking.] Leaf bud [and fruits] with very conspicuous villous hairs (1—)2—3 mm. [Inflorescences sessile.].
14'Twigs 6 mm diam. or less. Leaf bud with hairs 1.5 mm or less.
15Leaves pubescent below. Fruits 2.5-4.5 cm long.
15'Leaves glabrous below. Fruits 5 cm long or more.
16Inflorescences (best to be seen in male specimens) branched and with distinct ± flattened main peduncle, with flowers in subumbels or spikes. Fruits 2.5-4.5 cm long.
16'Inflorescences (sub)sessile scar-covered short-shoots lasting several flowering seasons. Fruits densely pubescent, 2.5-3.5 cm long. [Leaf bud with hairs up to 0.5(-l) mm. — Inland species; not from limestone.].
17Twigs 3-5(-10) mni diam. Leaves (12—) 15—35 cm long; lateral nerves 15—19(—25) per side. Leaf bud with hairs to 1(—1.5) mm. Buds rounded towards apex. Fruits 2.5-4.5 cm long, densely shaggy pubescent with hairs 0.5-1 mm. — Mainly a coastal species, but sometimes inland in degraded forest.
17'Twigs 1.5-3.5 mm diam. Leaves 11-20 cm long; lateral nerves 10-18 per side. Leaf bud with appressed hairs 0.5-1 mm. Buds sharply angular towards apex. Fruits 3.5-4 cm long, glabrescent or with thin scattered pale brown appressed hairs to 0.5 mm. — W Sarawak, NE Kalimantan; mostly from limestone.
18Twigs 2-3 mm diam. Leaves thinly coriaceous, base rounded or short acute; nerves not distinct above. Fruits 2.5-3.5 cm long, with scurfy hairs 0.2-0.3(-0.5) mm.
18'Twigs 3-4(-5) mm diam. Leaves coriaceous, base rounded or emarginate, sometimes short-acute; nerves distinct above. Fruits 2.5-3.5 cm long, with woolly hairs 0.5-1 mm.
19Male flowers: androecium ± sessile, with a collar of conspicuous hairs at base, sterile apex ± pubescent.
19'Androecium with distinct androphore, sterile apex without hairs. One collection, see note 3 under.
20Lower leaf surface concolorous with upper surface, not papillose. Fruits glabrous (early glabrescent).
20'Lower leaf surface distinctly papillose, generally paler than upper leaf surface. Fruits either with persistent indumentum, late glabrescent, or early glabrescent in M. gi-gantea.
21Leaves membranous, up to 24 cm long, base attenuate. Twigs up to 4 mm diam., smooth or striate, the bark of the twigs lower down cracking or not; leaf bud slender, 7-12 by 1-2 mm. Fruits to 8 cm long, pericarp 5-10 mm thick. — Stout forms, mainly from NE Borneo.
21'Leaves chartaceous, 15-35 cm long, base broadly rounded. Twigs (4-)5-6 mm diam., bark early conspicuously cracking; leaf bud stout, 10-15 by 4-6 mm. Fruits 7.5-9.5 cm long, pericarp 15-20 mm thick. — NE Borneo.
22Twigs 1.5—2.5(—3) mm diam. Leaves 7—12(—15) cm long, chartaceous, apex acute, blunt, or rounded; lateral nerves 10—15(—18) per side. Fruits 5-8 cm long, (late) glabrescent, hairs scurfy, 0.2 mm.
22'Twigs 3-6 mm diam. Leaves 14-35 cm long, coriaceous, the apex acute-acuminate; lateral nerves 13-22 per side. Fruits (5-)6-7 cm long, with conspicuous persistent shaggy hairs 1-2 mm.