KEY TO THE SPECIES

1Fertile pinnae similar to sterile pinnae
1'Fertile pinnae distinctly contracted
2Sori elliptic, in a single row between the veins, one in each main areole; rhizome scales without thickened midrib
2'Sori round, in many irregular rows between the veins, many in each main areole, rhizome scales with thickened midrib
3Fronds pinnate, pinnae shortly stalked, nectaries on small appendages at the pinna bases
3'Fronds lobed to pinnatifid, pinnae sessile, nectaries on the lamina
4At least fertile fronds stalked, without dilated base, rhizome scales appressed
4'All fronds sessile, with a dilated base, rhizome scales spreading or squarrose
5Sterile sessile fronds present.
5'Sterile fronds, if present, stalked
6Fertile fronds completely fertile, sori round, in several rows between costa and margin, rhizome scales peltate
6'Fertile fronds at the base sterile, sori on narrowed apical parts, quadrangular or elongated, in a single row between costa and margin, rhizome scales pseudopel-tate
7Sori mostly forming a continuous coenosorus, but rarely interrupted.
7'Sori quadrangular or elongated, not a continuous coenosorus
8Sporangia glabrous
8'Sporangia with 1-6 hairs
9Fertile pinnae between the sori distinctly contracted to a narrow inconspicuous wing
9'Fertile pinnae not contracted between the sori
10Rhizome not fully covered by the scales, with conspicuous phyllopods, fronds more than 20 cm distant
10'Rhizome completely covered with scales, phyllopods inconspicuous, fronds less than 10 cm apart
11Sori linear, coenosoroid, rhizome scales without thickened midrib.
11'Sori separate, rhizome scales with a thickened midrib
12Rhizome long-creeping, fronds over 20 cm distant, the midrib of rhizome scales black, sori round
12'Rhizome short-creeping, fronds less than 10 cm apart, midrib of rhizome scales brown
13Sori in several rows, confluent into irregularly quadrangular patches, abscission vein glabrous
13'Sori forming a single row of regular, quadrangular, large patches, abscission vein with short hairs