1 | Leaves paripinnate, without pseudogemmula or its scar |
1' | Leaves imparipinnate or pseudogemmulate |
2 | Inflorescences borne on leaves. — New Guinea |
2' | Inflorescences axillary, supra-axillary, ramiflorous or borne on bosses on bole. |
3 | Leaves ± densely pubescent, petiolules 3–6 mm long |
3' | Leaves sparsely hairy to glabrous, petiolules (5–)10–23 mm long |
4 | Inflorescences borne on long-lived bosses on bole. — Borneo |
4' | Inflorescences otherwise |
5 | Leaflets strongly asymmetric, petals 5 or 6, red, anthers 8–10 (limestone) |
5' | Leaflets not so, petals (3) 4 (5), white, anthers 6–9 (mountains) |
6 | Pseudopedicel c. 10 mm long, calyx with conspicuous annular thickening, petals 6-10 in 2 whorls — Malay Peninsula |
6' | Pseudopedicel, if present, much smaller |
7 | Calyx (10–)13–20(–23) mm long. — Borneo, Philippines |
7' | Calyx smaller 9 |
8 | Leaflets ± densely fulvescent abaxially, inflorescence to 30 cm, petals 9–14, anthers 15–20, stylehead discoid, seeds sarcotestal |
8' | Leaflets not so, inflorescence to 220 cm, petals 6–10, anthers 16–30, stylehead capitate, seeds arillate |
9 | Leaves imparipinnate |
9' | Leaves pseudogemmulate |
10 | Leaflets subglabrous abaxially — Sumatra, ?Java |
10' | Leaflets golden- to brown-pubescent, pilose or strigose adaxially |
11 | Anthers 3–5, corolla clavate in bud. — Malay Peninsula, Riau Arch |
11' | Anthers 6 or more |
12 | Petals 16 mm long. — Borneo |
12' | Petals 20 mm or longer |
13 | Leaflets rugose, surface strongly reticulate-areolate abaxially, shiny and glabrous adaxially save brown-tomentose midrib; seed sarcotestal. — Malay Peninsula |
13' | Leaflets smooth, abaxial surface not strongly reticulate-areolate; seeds arillate or unknown. — Borneo |
14 | Leaves strigose with ‘tinkling’ hairs, petals glabrous |
14' | Leaves appressed hirsute abaxially, glabrous or sparsely pubescent on veins adaxially, petals densely pubescent without. |
15 | Petals 26–37 mm long, pachycaul treelets with irritant fruit hairs |
15' | Petals smaller, fruit unarmed |
16 | Anther connective glabrous, seeds sarcotestal. — Malay Peninsula |
16' | Anther connective hairy, seeds arillate. — Borneo |
17 | One or more petals narrower than and enclosed by the others |
17' | Petals ± the same width or, at least, none completely enclosed by the others |
18 | Petals c. 8 mm long. — Sumatra |
18' | Petals 14 mm or longer |
19 | Costae c. 5–8 on each side of midrib, leaves to 38 cm. — Malay Peninsula |
19' | Costae c. 15 on each side of midrib, leaves to 150 cm. — Borneo, Celebes |
20 | Tube not conspicuously lobed or strongly crenulate (if unclear, follow the alternative) |
20' | Tube conspicuously lobed |
21 | Calyx 5–6.5 mm. — Celebes, New Guinea |
21' | Calyx up to 4 mm tall |
22 | Calyx 5–6.5 mm, shallowly cupulate. — Papua New Guinea |
22' | Calyx 5 mm, cylindrical-cupulate. — Celebes |
23 | Tube villous or sericeous without |
23' | Tube ± glabrous or at most sparsely hairy in middle or distal half without |
24 | Anthers 8. — Philippines |
24' | Anthers (3) 4–6 |
25 | Petals 5. — Sumatra |
25' | Petals (3) 4 |
26 | Petals glabrous, leaflets tomentose abaxially. — Celebes |
26' | Petals ± hirsute without 27 |
27 | Petals (3) 4, 7 mm long, ovary 4-locular. — New Guinea |
27' | Petals 4, much longer 28 |
28 | Ovary bilocular, seeds 2, arillate |
28' | Ovary 5-locular, seeds exarillate |
29 | Inflorescence villous, to 80 cm, calyx 1.5–2 mm long, anthers 8 or 9, petiolules c. 12 mm. — New Britain |
29' | Not this combination of characters |
30 | Flowerbuds less than 1.5 mm diam. — New Guinea |
30' | Flowers larger |
31 | Leaves with conspicuous venation on both sides (dry), coriaceous, glabrescent; petals c. 13.5 mm long, fruit rostrate |
31' | Leaves different, fruit not rostrate |
32 | Petals c. 12 mm long, flowers crowded towards distal end of inflorescence, fruit spherical |
32' | Petals up to 10 mm long, or, if slightly longer, then the tube villous within and the leaves pilose abaxially |
33 | Flowerbuds 9–10 mm, anthers 4–6 (7), style to 0.15 mm |
33' | Flowerbuds c. 11.5 mm, anthers 6–8, style 0.25 mm diam.. |
34 | Inflorescence borne on supra-axillary branch resembling supra-axillary inflorescence; petiolules fulvous tomentose, disk 1 mm |
34' | Inflorescences different; disk absent . |
35 | Inflorescences borne on supra-axillary branch resembling supra-axillary inflorescence; petiolules fulvous tomentose . |
35' | Inflorescences different. |
36 | Hairs simple. |
36' | Hairs stellate, disk present, seeds sarcotestal. — Borneo eastwards. |
37 | Corolla aestivation imbricate. |
37' | Corolla aestivation valvate, seeds sarcotestal |
38 | Costae 15–22 on each side of leaflet midrib, flowerbuds clavate, anthers 3–5, pachycaul treelet. — Malay Peninsula, Riau |
38' | Not this combination of characters |
39 | Ovary 4-locular, seeds (3) 4. |
39' | Ovary 2-locular, seeds 2. |
40 | Disk obscure. |
40' | Disk prominent, subtubular to 1 mm. |
41 | Calyx 4-5-lobed, petals 5 or 6, anthers scarcely locellate . |
41' | Calyx margin entire, petals (3) 4 (5), anthers locellate |
42 | Disk present. |
42' | Disk absent |
43 | Venation very prominent on both sides (dried), seeds 3 cm diam. — Borneo |
43' | Venation and seeds different |
44 | Petals 18 mm long. — Philippines |
44' | Petals shorter . |
45 | Tube crenate, anthers hairy . |
45' | Tube lobes long-triangular, anthers glabrous . |
46 | Disk present |
46' | Disk absent |
47 | Twigs subglabrous 48 |
47' | Twigs fawn pubescent. |
48 | Petals 13–19 by 2 mm, tube lobes ± truncate, flowers in panicles, fruit 4.5 cm or more . |
48' | Petals 8–12(–19) mm, tube lobes ± laciniate, flowers in panicles only if flowers small; fruit to 2.1 cm diam . |
49 | Petals 16 mm long. — Philippines . |
49' | Petals 9–13 mm long |
50 | Flowers 11–12 mm long. — Borneo, Celebes |
50' | Flowers less than 8 mm long . |
51 | Inflorescence velutinous, style glabrous. — New Guinea |
51' | Inflorescence subglabrous, style densely hairy |