KEY TO THE SPECIES

1Hairs on upper surface of rachis ctenitoid, shrivelled when dried or with obtuse apical cells
1'Hairs on upper surface of rachis persistently terete with acute apical cells
2Sori exindusiate; scales less than 2 mm long
2'Sori indusiate; scales much longer
3Frond wholly anadromous
3'Frond catadromous distally