KEY TO THE SUBSPECIES

1Inflorescence and flowers densely ferrugineous villous pubescent. Often at high altitudes
1'Inflorescence and flowers sparsely to densely grey or brown appressed pubescent. Lowlands.
2Primary leaf veins 16-26 pairs; mature leaves 9-15.7 cm long, oblong (index 2.3-3.65). Fruit exocarp usually densely verrucose
2'Primary leaf veins 10-16 pairs; mature leaves 5-10.5 cm long, elliptic (index 1.7-2.7), rarely oblong. Fruit exocarp usually sparsely verrucose