| 1 | Leaflets nearly always equal-sided; midrib and nerves nearly always (distinctly) sunken above; petiolules often grooved |
| 1' | Leaflets mostly distinctly oblique at base; midrib nearly always flat or prominulous above, nerves above prominulous; petiolules rarely grooved |
| 2 | Fruits long-aculeate. |
| 2' | Fruits smooth to warty. |
