Leaves opposite, those of a pair equal, entire, petiolate; Inflorescence of axillary and, rarely, terminal cymes, often very short, appearing fasciculate; Flowers 5-merous, actinomorphic. Sepals connate at the base; Stamen insertion variable, completely included within the tube, free from the style head; Fruit brown or yellowish, a fleshy berry, spherical, ellipsoid or pear-shaped, indehiscent; Seeds compressed ovoid, without a coma, smooth, with a very thin endosperm and thick horny cotyledons.
Asia-Tropical, western Malesia present
16 species in Indochina and western Malesia; in Malesia 15 species.
2 Merrill wrote the names of several ‘new species’ on herbarium labels which he never actually published. 1 There is some question as to the assignment to species of sterile specimens with long narrow leaves, see Middleton (1993) op. cit.
Backer & Bakh.f. - in Fl. Java. 1965: 224
Pit. - in Lecomte, Fl. Indo-Chine 3. 1933: 1094
D.J. Middleton - in Blumea. 1993: 2
D.J. Middleton - in Fl. Thailand. 1999: 18
Benth. & Hook.f. - in Gen. Pl. 1873: 691
G. Don - in Gen. Hist. 1837: 101
Backer & Bakh.f. - in Fl. Java. 1965: 223
Roxb. - in Tree Fl. Sabah & Sarawak. 2004: 6
Kuntze - in Revis. Gen. Pl. 1891: 412
Pichon - in Mém. Mus. Natl. Hist. Nat. 1948: 153
Pierre, Bull. Mens. Soc. Linn. Paris. 1898: 94
A.DC. - in Prodr. 1844: 321
Coode et al., Checklist Pl. Brunei. 1996: 29
Pichon - in Mém. Mus. Natl. Hist. Nat. 1948: 154
Hook.f. - in Fl. Brit. India. 1882: 623
K. Schum. - in Engl. & Prantl, Nat. Pflanzenfam. 4. 1895: 130