Willughbeia
Content
Description
Leaves opposite, those of a pair equal, entire, petiolate;
Inflorescence of axillary and, rarely, terminal cymes, often very short, appearing fasciculate;
Flowers 5-merous, actinomorphic.
Sepals connate at the base;
Stamen insertion variable, completely included within the tube, free from the style head;
Fruit brown or yellowish, a fleshy berry, spherical, ellipsoid or pear-shaped, indehiscent;
Seeds compressed ovoid, without a coma, smooth, with a very thin endosperm and thick horny cotyledons.
Distribution
Asia-Tropical, western Malesia present
16 species in Indochina and western Malesia; in Malesia 15 species.
Taxonomy
2 Merrill wrote the names of several ‘new species’ on herbarium labels which he never actually published.
1 There is some question as to the assignment to species of sterile specimens with long narrow leaves, see Middleton (1993) op. cit.
Citation
Backer & Bakh.f. 1965 – In: Fl. Java: 224
Pit. 1933 – In: Lecomte, Fl. Indo-Chine 3: 1094
D.J. Middleton 1993 – In: Blumea: 2
D.J. Middleton 1999 – In: Fl. Thailand: 18
Benth. & Hook.f. 1873 – In: Gen. Pl.: 691
G. Don 1837 – In: Gen. Hist.: 101
Backer & Bakh.f. 1965 – In: Fl. Java: 223
Roxb. 2004 – In: Tree Fl. Sabah & Sarawak: 6
Kuntze 1891 – In: Revis. Gen. Pl.: 412
Pichon 1948 – In: Mém. Mus. Natl. Hist. Nat.: 153
Pierre 1898: Bull. Mens. Soc. Linn. Paris: 94
A.DC. 1844 – In: Prodr.: 321
Coode et al. 1996: Checklist Pl. Brunei: 29
Pichon 1948 – In: Mém. Mus. Natl. Hist. Nat.: 154
Hook.f. 1882 – In: Fl. Brit. India: 623
K. Schum. 1895 – In: Engl. & Prantl, Nat. Pflanzenfam. 4: 130