KEY TO THE SPECIES

1Basal nerves running up throughout the length of the leaf or nearly so.
1'Basal nerves running up to ½-⅔ the length of the leaf.
2Leaf thick-coriaceous, lower surface densely set with soft erect hairs; margin distinctly serrate; reticulations prominent beneath. Inflorescence ♂ or ♀, at anthesis condensed and shorter than the petiole
2'Leaf chartaceous to thin-coriaceous, lower surface glabrous or sparsely appressed-hairy; margin finely serrulate to subentire; reticulations obscure. Inflorescence ♂ or ♂♀, at anthesis lax and longer than the petiole
3Leaf thick-coriaceous, upper surface strongly rugose and scabrous, lower surface densely pubescent; midrib, nerves, and reticulations prominent beneath
3'Leaf chartaceous to thin-coriaceous, upper surface not or hardly rugose nor scabrous, lower surface sparsely appressed pubescent or glabrous; midrib, nerves, and reticulations only slightly raised beneath.
4Leaf elliptic-lanceolate, more or less glabrous; nerves more than 4 on each side, straight and ascending at a narrow angle (less than 40°) from the midrib. Inflorescences mostly ♂♀, very rarely ♂ or ♀
4'Leaves ovate to ovate-elliptic, underneath sparsely appressed pubescent; nerves less than 4, usually 3 on each side, arcuating at a wider angle (c. 45-60°) from the midrib. Inflorescences ♂ or ♀, rarely ♂♀