Asia-Tropical: Bismarck Archipelago (Bismarck Archipelago present); New Guinea present
Malesia: New Guinea, Bismarck Archipelago.
1 This variety occurs generally at lower altitudes than the type variety, but more or less transitional forms occur in the range of overlap. The discontinuity between the two varieties may be due to the absence of suitable habitats in the intermediate range. 2. Plants are occasionally infested with a gall-inducing sawfly, which causes the sori to swell. The formation of sporangia is suppressed in favour of paraphyses. This results in the formation of characteristically protruding tufts.