Guioa bijuga

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Guioa bijuga


Tree, 3-30 m high, dbh 5 cm to 1 m; Branchlets usually glabrous, at most shortly sericeous when young, hairs brownish; Leaves 1-4-ju-gate; Inflorescences axillary (to pseudoterminal), (un-branched to) branching basally to especially along the terete to flattened, glabrous to pilose, 0.7-16.5 cm long axis; Flowers 3.5-4.2 mm in diam. Sepals 5, ovate, margin pilose, with glands, outside and inside (sub)glabrous, green; Petals 5, obovate, 1.8-3.8 by 0.7-1.7 mm, white to yellow, blade obovate, gradually decurrent into the 0.4-1.2 mm high claw, margin pilose, outside and inside (sub)glabrous, apex emarginate to acute; Stamens 8; Fruits with 1-3 well developed lobes, 1.2-2.3 by 1.3-2.6 cm, smooth to somewhat ribbed or rugose-ruminate, glabrous, blackish when dry; Seeds (globose to) obovoid, 8-9.8 by 7-7.8 mm;


Asia-Tropical: Borneo present; Malaya (Peninsular Malaysia present); Philippines (Philippines present); Sumatera (Sumatera present); Thailand (Thailand present), Balabac I present, Palawan present
Thailand to Malesia: Peninsular Malaysia, Sumatra, Borneo, Philippines (Balabac I., Palawan).


1. Several specimens, growing on poor white sand, have smaller leaflets (3.2-14 by 1.5-7.9 cm), which are very thick and scleromorphic; leaflets of specimens on richer soil are 3.9-19.6 by 1.8-8.4 cm, thinner, less scleromorphic. The same phenomenon can be observed in G. acutifolia.
One specimen is exceptional; S 32186: leaflets with many pockets, lower surface very papillate and somewhat sericeous.
The difference between G. bijuga and G. pubescens is often vague on Borneo. Usually, G. bijuga has elliptic leaflets which are smooth and usually glabrous below; the fruits are glabrous too. Guioa pubescens usually has ovate leaflets, which are papillate below and sericeous, the fruits are glabrescent. On Borneo, G. bijuga can be papillate and somewhat sericeous, while G. pubescens can have rather elliptic and more glabrous leaflets, especially in Sarawak and on the Kinabalu. Even there the more ovate shape of the leaflets and the few hairs remaining on the fruit indicate G. pubescens and the flattened above to winged rachis indicates G. bijuga. There are also anatomical differences: G. bijuga always has secretory idioblasts, while G. pubescens always lacks them.


Masamune 1942: Enum. Phan. Born.: 427
Briinig 1968: Heidewald Sarawak Brunei: 372
Radlk. 1879 – In: Sitzungsber. Math.-Phys. CI. Konigl. Bayer. Akad. Wiss. Munchen: 521,611
Corner 1939 – In: Gard. Bull. Str. Settl.: 44
Radlk. 1940: Wayside Trees: 588
Ridley 1922 – In: Fl. Malay Penins.: 505
Corner 1978: Gard. Bull. Sing.: 153
Welzen 1989: p. 183. – In: Leiden Bot. Series: f. 68, 69
Radlk. 1933 – In: Engl., Pflanzenr. 98: 1165
Yap 1989 – In: Tree Fl. Malaya: 441
Meijer 1967 – In: Bot. News Bull.: 74
Radlk. 1933 – In: Engl., Pflanzenr. 98: 1177