Echinodorus macrophyllus subsp. scaber Echinodorus macrophyllus subsp. scaber (Rataj) R.R. Haynes & Holm-Niels. in Brittonia 38: 331. 1986 sec. B. Ståhl & al. 2009≡Echinodorus scaber Rataj in Folia Geobot. Phytotax. 4: 438. 1969Holotype: Guyana, Canje R., Jenman 4310