5 — KEY TO THE SPECIES OF BORNEO (25 species)

1Leaf blade foliolate (in T. wawrae sometimes, in T. elmeri rarely, partly or all simple)
1'Leaf blade simple (unlobed or lobed)
2Leaf blade 3-foliolate, membranous or chartaceous; probract membranous, flat (sometimes caducous). Tendrils simple or 2-branched. Male rachis 1-1.5 mm thick. Fruit 6-9 cm long
2'Leaf blade 3-5-foliolate, chartaceous or coriaceous; probract chartaceous, concave. Tendrils 2- or 3-branched. Male rachis 2-3 mm thick. Fruit more than 10 cm long
3Plant (sub)annual; monoecious. [Leaf blade membranous, green-yellow on drying.] Corolla 20(-30) mm diameter. Male bracts 1(-2) mm long, often caducous. [Fruit ovoid, or long-cylindrical in cultivated var. anguina.] Seeds with undulate margin
3'Plant (sub)perennial; monoecious or dioecious. Corolla usually more than 30 mm diam. (smaller in T. postari). Male bract 3 mm long or more. Seeds tumid or compressed, not with undulate margin
4Whole plant conspicuously (densely) soft hairy; hairs 1 mm long or more
4'Whole plant glabrous or (sparsely) hairy; hairs less than 1 mm long
5Indumentum grey-rusty villose. Tendrils 5-15-branched, point of branching 1-2 cm from the base. Leaf blade without conspicuous mucro. Fruit broad ellipsoid (-globose), 9-15 cm long. Seeds compressed
5'Indumentum grey, hirsute. Tendrils 3-5-branched, point of branching 2-4 cm from the base. Leaf blade with conspicuous mucro, 13-25 mm long. Fruit ellipsoid, 9-10 cm long. Seeds tumid
6Leaf blade at transition to petiole with 2 gland-bearing auricles. [Male bracts linear. Fruit in a raceme, globose.] Seeds subcircular with broad radially striate margin
6'Leaf blade at base without auricles. Seeds tumid or compressed, without radially striate margin
7Flowers conspicuously brown hairy (fruit and seed not known)
7'Flowers greyish hairy or glabrous
8Inflorescences borne on leafy twigs
8'Inflorescences borne on older stems
9Stem sharply angular or ridged
9'Stem striate or grooved
10Stem and leaf blade brown on drying, reddish tinged. Leaf apex without acumen. Probract narrowly elliptic. Male raceme solitary. Seeds compressed, with ± cuneate base, and broadly rounded apex
10'Stem and leaf blade green on drying, not reddish tinged. Leaf apex often with a 10-25 mm long acumen. Probract absent. Male (and female) racemes mostly fascicled, pendent. Seeds tumid
11Plant with fruit (and/or female flowers). Fruit of T. longispicata not known
11'Plant with male flowers
12Seeds tumid
12'Seeds compressed
13Leaf venation ± flat beneath, usually minutely hairy. Fruit pulp not fibrous. Seeds barrel-shaped or (sub)fusiform (Fig. 91b)
13'Leaf venation much raised beneath, glabrous or hairy. Fruit pulp coarsely fibrous. Seeds butterfly-shaped
14Plant free-climbing
14'Plant growing adherent to tree trunks. (Fruit not known)
15Fruit globose
15'Fruit ovoid or ellipsoid
16Fruiting pedicel with two parts of different surface and colour
16'Fruiting pedicel evenly coloured
17Fruiting pedicel less than 10 mm thick
17'Fruiting pedicel 10-15 mm thick
18Leaf blade ± 5-angular, blackish on drying; glands several, 0.5-1 mm diam., close to the insertion of the petiole. Seeds 9-14 by 4-5 mm, chisel-shaped pointed at base
18'Leaf blade unlobed or (deeply) lobed, brown-blackish on drying; glands absent or few, 1-2 mm diam., scattered but mainly towards the base of the blade. Seeds 18-22 by 7-8 mm, rounded at both ends
19Leaf blade pale green on drying. Tendrils simple
19'Leaf blade green or brown on drying. Tendrils (usually) 2-4(-5)-branched
20Leaf blade unlobed or shallowly 3-lobed, green on drying. Fruiting pedicel c. 4 cm long. Seeds 13-18 by 7-9 mm
20'Leaf blade deeply 5-7-lobed, greenish brown or blackish on drying. Fruiting pedicel c. 3 cm long. Seeds 17-20 mm long
21Plant slender, leafy stem 2-3 mm diameter. Pericarp ± leathery, thin
21'Plant stouter, leafy stem 3-4 mm diameter or more. Pericarp carnose, c. 5 mm thick when dry. [Seeds c. 10 mm long. Leaf blade 3- or 5-lobed.]
22Leaf blade ovate-hastate, usually ± lobed; glands c. 0.5 mm diameter. Fruit 6-10 cm long, yellow striped. Seeds 15-18 mm long 43. T. wawrae and
22'Leaf blade (narrowly) ovate, unlobed; glands 1-2 mm diameter. Fruit c. 5 cm long (ripening colour not known). Seeds c. 13 mm long
23Fruit (possibly) ripening green. — Peat swamp forest
23'Fruit ripening red
24Leaf blade (±) hairy, 3-5-lobed, margin (coarsely) dentate. Young parts of shoots reddish brown on drying. Probract narrowly elliptic or linear, 10-20 mm long. Petal fringes tinged pinkish. Fruiting pedicel (1.5-)2-5 cm long
24'Leaf blade glabrous or early glabrescent, 3-lobed to 1/4-1/2 deep, margin entire or dentate. Young parts of shoots brown on drying. Probract obovate or elliptic, 5(-10) mm long. Petals fringes white. Fruiting pedicel 0.5-2.5 cm long
25Rachis of male raceme 1-1.5(-2) mm thick. Leaf blade frequently unlobed
25'Rachis of male raceme 2-10 mm thick. Leaf blade mostly lobed
26Blade glands c. 2 mm diameter
26'Blade glands less than 1 mm diameter, or absent
27Leaf blade glabrous beneath (hairy in forma hirsuta). Persistent part of pedicel almost absent (sometimes persistent)
27'Leaf blade ± hairy beneath. Persistent part of pedicel 2 mm long or more
28Venation on lower leaf surface raised. Peduncle short, 0.5-3 cm long
28'Venation on lower leaf surface ± flat. Peduncle longer, (2-)5-8(-17) cm long
29Leaf blade ± 5-angular. Bracts entire. Sepals mostly with narrow sidelobes
29'Leaf blade various. Bracts incised. Sepals entire or lobed
30Persistent part of pedicel 5-10 mm long. Bracts 7-12 mm long
30'Persistent part of pedicel shorter or absent. Bracts usually larger
31Rachis 2-4 mm thick
31'Rachis 5-10(-20) mm thick
32Rachis ± zigzag
32'Rachis straight
33Bracts less than 15 mm long
33'Bracts more than 15 mm long
34Bracts broad, deeply fan-shaped incised. Plant dull blackish on drying
34'Bracts incised, but not fan-shaped. Plant green or (reddish) brown on drying
35Leaf blade ± hairy. Shoots reddish tinged. Petal fringes tinged pinkish
35'Leaf blade glabrous or early glabrescent. Shoots green or brown on drying. Petal fringes white
36Leaf blade deeply 5-7-lobed. Blade glands few (or absent), near the insertion of the petiole, 1-3 mm diameter. Probract narrowly elliptic, without glands
36'Leaf blade 3-lobed to 1/4-1/3 of the blade. Blade glands scattered, less than 1 mm diameter. Probract ± (ob)ovate or elliptic, with conspicuous glands
37Bracts entire, pale yellow soft hairy. Tendrils simple
37'Bracts incised. Tendrils 2-4-branched
38Leaf blade unlobed or shallowly (2- or) 3-lobed. — Lowland
38'Leaf blade deeply 5(-7)-lobed. — Lowland and montane