Primary tabs



Trees, dioecious. Leaves simple, distichous; Stipules falcate, early caducous, tomentose. Inflorescences dense axillary clusters of 2-many flowers (reduced thyrses), tomentose, brachyblasts increasing in size with age; Flowers actinomorphic; Fruits globose rhegmas, outside densely tomentose, reticulately wrinkled when dry, with 6 raised lines, dehiscing completely septicidally and partly loculicidally into 3 bifid pieces; Seeds ovoid, flattened, 1-3 per fruit, black, glossy, abaxially covered up to ⅔ rd by a thin, sometimes lobed aril, lobes sometimes touching adaxially;


Asia-Tropical: Borneo present (Sarawak present); Malaya (Peninsular Malaysia present, Singapore absent); Philippines (Philippines present); Sulawesi (Sulawesi present); Sumatera (Sumatera present), W Malesia present
The genus comprises 3 species, all found in Malesia. One is widespread in W Malesia (Peninsular Malaysia, Sumatra, Borneo, Sulawesi; not known from Singapore), the others are endemics in Sarawak or in the Philippines.


Whitmore - in Tree Fl. Malaya. 1973: 134
G.L.Webster - in Ann. Missouri Bot. Gard. 1994: 65
Welzen, Bulalacao & On - in Blumea. 1995: 368
Hook.f. - in Kew Bull. 1981: 350
Bulalacao & On - in Natl. Mus. Pap. (Philippines). 1992: 67
Hook.f., Alphabet. Enum. Euphorb. Philipp. Isl. 1983: 46
Hook.f. - in Pflanzenr. 1919: 42
Pax & K.Hoffm. - in Engl., Pflanzenr. 147. 1911: 95
Radcl.-Sm., Gen. Euphorbicearum. 2001: 115
Ridl. - in Fl. Malay Penins. 1922: 290
Airy Shaw - in Kew Bull. 1975: 201
Pax - in Engl. & Prantl, Pflanzenfam. 3. 1890: 84