Understory treelets or medium to large trees. Stipules wanting. Leaves alternate, pinnately compound, with a rudimentary distal leaflet; leaflets opposite or alternate, with entire margins; petiolules usually swollen. Inflorescences axillary, distal or seldom cauliflorous, paniculate or racemiform thyrses, with flowers in lateral, simple or compound dichasia. Flowers actinomorphic, functionally staminate or pistillate (plants dichogamous); sepals 5, connate at base, usually shorter than the petals; petals 5, distinct, erect or reflexedwith a simple or bifid petaloid appendage; disc annular; stamens 5-8, the filaments of unequal length, free, villose or glabrous, the anthers basifixed; ovary 3-locular, each locule with a single ovule. Fruits ellipsoid or globose, with leathery to woody pericarp. Seed usually one or less often two or three per fruit, with a fleshy, usually edible testa.