1Bases of pinnae, at least on their acroscopic side, cordate
1'Bases of pinnae hastate, their basal auricles acuminate
2Pinnae of mature plants 10 cm or more long
2'Pinnae of mature plants c. 3.5 cm long
3Few pairs of lower pinnae slightly reduced and more widely spaced; scales bearing marginal teeth
3'Many pairs of lower pinnae gradually much reduced; scales lacking marginal teeth
4Sori exindusiate; acroscopic bases of middle pinnae not cordate
4'Sori indusiate; acroscopic bases of middle pinnae cordate