Meliosma pinnata subsp. arnottiana
Content
Description
Small to medium-sized, rarely big tree, up to c. 20(-30) m.
Leaves (2-)3-7(-8)-jugate;
Sepals glabrous or the outer ones rarely with a few hairs.
Petals glabrous, inner ones sometimes a bit ciliolate or frayed at the tips of the lobes.
Panicles erect, spreading, lax to dense, lower primary side-axes usually subtended by small or reduced leaves.
Ovary densely pubescent, very rarely subglabrous.
Distribution
Asia-Temperate: Taiwan (Taiwan present), Asia-Tropical: Malaya (Peninsular Malaysia present); Philippines (Philippines present); Sri Lanka (Sri Lanka present), Batan Is present, Japan present, Karo present, Luzon present, Mindanao present, N. Sumatra present, S. Korea present, SE. Asia present, W. & Central Java present, W. Malesia present
Sri Lanka throughout SE. Asia to China, S. Korea, Japan and Taiwan; in Malesia: N. Sumatra (Karo), Malay Peninsula, W. & Central Java, Philippines (Batan Is., Luzon, Mindanao). Rare in W. Malesia. .
Notes
Attention should be given to the relation between subsp. arnottiana and subsp. pendula in the Philippines (for the relation to subsp. macrophylla, see under that subspecies). In the mountains of Luzon both subspecies have been collected, subsp. pendula above 1800 m altitude and subsp. arnottiana from c. 800-900 m up to c. 2400 m. Locally, e.g. on Mt Santo Thomas, they have been found together, but doubtless intermediate specimens are not observed. It is possible that in such localities these subspecies mutually behave as species; population studies in the field might yield more evidence with regard to this.
The same problem arises in W. Malesia, where subsp. arnottiana has been collected (rarely). In Sumatra and in Java its relation to subsp. ferruginea is interesting since there is an altitudinal zone of overlap between both, though subsp. ferruginea generally occurs lower than subsp. arnottiana. In Java the situation is as follows: subsp. ferruginea is by far the most common of both, subsp. arnottiana having only been collected on a few mountains. Of these it is only G. Salak and G. Gedeh where both subspecies have been found. Only of G. Gedeh more detailed ecological evidence is available: KOORDERS () stated that subsp. ferruginea occurs at c. 1400 m altitude and that subsp. arnottiana ('M. glau-ca') occupies a zone between 1800 and 2400 m, being especially abundant at c. 2200 m. This does suggest the existence of ecological differentiation, but since subsp. arnottiana on other mountains also grows at lower altitudes, the situation remains unclear.
The same problem arises in W. Malesia, where subsp. arnottiana has been collected (rarely). In Sumatra and in Java its relation to subsp. ferruginea is interesting since there is an altitudinal zone of overlap between both, though subsp. ferruginea generally occurs lower than subsp. arnottiana. In Java the situation is as follows: subsp. ferruginea is by far the most common of both, subsp. arnottiana having only been collected on a few mountains. Of these it is only G. Salak and G. Gedeh where both subspecies have been found. Only of G. Gedeh more detailed ecological evidence is available: KOORDERS () stated that subsp. ferruginea occurs at c. 1400 m altitude and that subsp. arnottiana ('M. glau-ca') occupies a zone between 1800 and 2400 m, being especially abundant at c. 2200 m. This does suggest the existence of ecological differentiation, but since subsp. arnottiana on other mountains also grows at lower altitudes, the situation remains unclear.
Citation
GAMBLE 1918 – In: Fl. Pres. Madras: 256
K. & V. 1903 – In: Bijdr.: 137
Blume 1923 – In: Fl. Tjibodas: 157
WALP. 1852 – In: Ann.: 225
Merr. 1923 – In: Enum. Philip. Fl. Pl.: 517
Miq. 1859 – In: Fl. Ind. Bat.: 615
K. & V. 1903 – In: Bijdr.: 137
WALP. 1842 – In: Rep.: 416, 423
Elmer 1908 – In: Leafl. Philip. Bot.: 492
WIGHT 1840 – In: Ill. Ind. Bot.: 142
Koord. 1912 – In: Exk. Fl. Java: 546
TRIM. 1893 – In: Fl. Ceyl.: 315
HALL.Ƒ 1910 – In: Meded. Rijksherb.: 2
HASSK. 1858 – In: Hort. Bog.: 140
HALL.f. 1910 – In: Meded. Rijksherb.: 2
Merr. 1923 – In: Enum. Philip. Fl. Pl.: 517
Blume 1871: Illustr.: 74
BEDD. 1871 – In: Fl. Sylv.: 77
WALP. 1852 – In: Ann.: 225
K. & V. 1903 – In: Bijdr.: 135
THW. 1858: Enum. Pl. Zeyl.: 59
Miq. 1859 – In: Fl. Ind. Bat.: 615
BRANDIS 1906: Indian Trees: 195
BAKERƑ 1924 – In: Rendle, J. Bot. 62: Suppl. 30
Koord. 1912 – In: Exk. Fl. Java: 546
Hook.f. 1876 – In: Fl. Brit. India: 6
BEDD. 1871: p. 77. – In: Fl. Sylv.: t. 160