Goniophlebium
Distribution
Asia-Tropical, Australasia: Queensland (Queensland present), Northern India to Japan present, Pacific: Samoa (Samoa present)
Northern India to Japan, Samoa, Australia (Queensland). Throughout Malesia.
Taxonomy
Goniophlebium has been regarded as a part of Polypodium, closely allied to a number of pinnate species from the Neotropics. Several authors considering Goniophlebium distinct from Polypodium have included the Neotropic as well as the Asian species. However, Copeland (e.g., 1929, 1947) pointed out that the Asian and American lines in this group should be considered distinct, and circumscribed the Asian Goniophlebium species by the separate, articulate pinnae. In this he was not widely followed, and many authors continued to use the name Polypodium for the Asian representatives. Copeland’s (l.c.) distinction is followed here, but Gonio- phlebium is distinguished by a combination of characters (see also Rödl-Linder 1990): rhizome with sclerenchyma strands, rhizome scales pseudopeltate, clathrate, lamina pinnate or pinnatifid, venation simple, with at most one, excurrent included vein in each areole.
Within Goniophlebium, Rödl-Linder distinguished five groups. The G. percussum-group is diagnosed by having all pinnae articulated. This group comprises all Malesian species except G. mengtzeense, G. prainii, G. pseudoconnatum and G. subauriculatum. The other groups are not clearly diagnosed. Goniophlebium pseudoconnatum and G. subauriculatum are placed close together, with a number of mainly Asian species, of which G. prainii occupies a somewhat uncertain, basal position in the group. Goniophlebium mengtzeenze is considered most closely related to the Asian G. argutum.
Within Goniophlebium, Rödl-Linder distinguished five groups. The G. percussum-group is diagnosed by having all pinnae articulated. This group comprises all Malesian species except G. mengtzeense, G. prainii, G. pseudoconnatum and G. subauriculatum. The other groups are not clearly diagnosed. Goniophlebium pseudoconnatum and G. subauriculatum are placed close together, with a number of mainly Asian species, of which G. prainii occupies a somewhat uncertain, basal position in the group. Goniophlebium mengtzeenze is considered most closely related to the Asian G. argutum.
Citation
Rödl-Linder 1990 – In: Blumea: 371
Copel. 1929 – In: Univ. Calif. Publ. Bot.: 109
Pichi Serm. 1973 – In: Webbia: 468
J. Sm. 1875: Hist. Fil.: 92
C. Presl 1883: Handb. Ferns Brit. India: 316
Farwell 1931 – In: Amer. Midl. Nat.: 295
C. Presl 1947: Gen. Fil.: 181
Bedd. 1866: Ferns Brit. India: 163, 206
Hennipman et al. 1990: p. 225. – In: Kramer & Green, Fam. & Genera Vasc. Pl. 1: (‘Goniophlebium Group’).
Zijlstra et al. 1987 – In: Taxon: 759
Fée 1850–1852: Gen. Filic.: 254