Milichiella argyrogaster
Description
Body length: 3.5 mm; wing length: 3.0 mm.
Male coloration and vestiture: frons subshiny black, ocellar triangle and orbital plates shiny black, lunule ?, basoflagellomere black, face ?, palpus black; thorax with mesonotum sligthly to normally brown microtomentose, scutellum and pleura brown microtomentose, wing hyaline, veins light brown, brown spot at tip of vein R1 absent, calypter hyaline with light brown margin, halter black, legs black except for brown basitarsi; abdomen with T2-5 silvery microtomentose.
Morphology. Head: frons with ? orbital and ? frontal setae, anterior frontal seta ?. In males frons ?x as wide at posterior margin than at base of antenna, arista about ?x as long as basoflagellomere wide, pubescence on arista ?, posterior eye margin with notch (7oc) and emargination (2oc).
Thorax: 2+1? dc, ? prsc, 1 pprn, 1+0.8 setae prs and row of shorter setae, ? keps setae, posterior prsc about as long as posterior dc, seta between posterior prsc and posterior dc present. Wing with vein R4+5 and M1 converging (2.4), and ultimate section of vein M1 2.2x as long as penultimate section. Fore femur strongly setulose, tibial organ absent.
Male abdomen: Posteromedial triangular projection of T1 into T2 present. Tergal chaetotaxy: T2 anterolateral corner with 1-2 lines of setae, T2-5 posterior margin with 1 row of setulae. S5 about as long as wide, ?x as long and 2.7x as wide as S4, nearly heartshaped, setulose. Male genitalia with apical seta on cercus as long as longest epandrial seta, 1 seta on subepandrial plate, surstylus about as long as epandrium high, bent posteriorly, dilated apically (2.6x), apex flat, 6.7x as long as wide, structure of lateral surface smooth, medial surface with line of setae near anterior margin.
Male coloration and vestiture: frons subshiny black, ocellar triangle and orbital plates shiny black, lunule ?, basoflagellomere black, face ?, palpus black; thorax with mesonotum sligthly to normally brown microtomentose, scutellum and pleura brown microtomentose, wing hyaline, veins light brown, brown spot at tip of vein R1 absent, calypter hyaline with light brown margin, halter black, legs black except for brown basitarsi; abdomen with T2-5 silvery microtomentose.
Morphology. Head: frons with ? orbital and ? frontal setae, anterior frontal seta ?. In males frons ?x as wide at posterior margin than at base of antenna, arista about ?x as long as basoflagellomere wide, pubescence on arista ?, posterior eye margin with notch (7oc) and emargination (2oc).
Thorax: 2+1? dc, ? prsc, 1 pprn, 1+0.8 setae prs and row of shorter setae, ? keps setae, posterior prsc about as long as posterior dc, seta between posterior prsc and posterior dc present. Wing with vein R4+5 and M1 converging (2.4), and ultimate section of vein M1 2.2x as long as penultimate section. Fore femur strongly setulose, tibial organ absent.
Male abdomen: Posteromedial triangular projection of T1 into T2 present. Tergal chaetotaxy: T2 anterolateral corner with 1-2 lines of setae, T2-5 posterior margin with 1 row of setulae. S5 about as long as wide, ?x as long and 2.7x as wide as S4, nearly heartshaped, setulose. Male genitalia with apical seta on cercus as long as longest epandrial seta, 1 seta on subepandrial plate, surstylus about as long as epandrium high, bent posteriorly, dilated apically (2.6x), apex flat, 6.7x as long as wide, structure of lateral surface smooth, medial surface with line of setae near anterior margin.
Diagnosis
Differs from congeners in the combination of 1 pprn, calypter hyaline with light brown margin, black legs, silvery microtomentose male T2-5, in the shape and shorter setae of the surstylus.
Discussion
There is one probable type in MNHNP. The specimen fits the description and appears to be very old, however, both the locality label as well as the identification label are written by Seguy. The male specimen is in relatively good condition, head deformed.
Distribution
Asia-Temperate: Russian Far East (Primorye present); Europe: Eastern Europe (North European Russia present); Middle Europe (Austria present, Czechoslovakia (Czech Republic present, Slovakia present), Hungary present); Southwestern Europe (France present)
Citation
catalog, Spain (Carles-Tolrá & Báez 2002: Milichiidae, p. 174. – In: Carles-Tolrá, Catálogo de los Diptera de España, Portugal y Andorra (Insecta) 8: 174)
Palaearctic catalog (Papp 1984: Family Milichiidae + Carnidae, pp. 110-123. – In: Soos, A. & Papp, L., Catalogue of Palaearctic Diptera 10, pp. 110-1: 113)
citation
figs of male genitalia, sternites and photo of male dorsal habitus, Japan (Iwasa 1999: The genus Milichiella Giglio-Tos (Diptera, Milichiidae) of Japan. – Med. Ent. Zool. 50: 27-32: 30 (as Milichiella argylogaster4))
Milichiella argyropasta1: Aldrich 1931: New acalyptrate Diptera from the Pacific and Oriental Regions. – Proc. Hawaii. ent. Soc. 7: 395-399: 398 (as Milichiella argyropasta1)
Palaearctic catalog (Becker 1905: Katalog der paläarktischen Dipteren 4: 237-240: 238 (as Stenoporomyia tiefii))
Hungary (Papp 1978: 72. család: Milichiidae + Carnidae. – Fauna Hung. 15: 9-50: 17)
larvae and pupae under wormeaten bark of piece of lime tree (Perris 1876: Nouvelle promenade entomologique. – Annls Soc. ent. Fr. (N. S.) (5)6: 208: 208 (as Milichia argyrogaster))
diagnosis (Becker 1907: Die Dipteren-Gruppe Milichiinae. – Annls hist.-nat. Mus. natn. hung. 5: 507-550: 534 (as Milichiella argrogastra))
biology, development, description of egg, larva and puparium (Rohácek, J. 1995: Habitat, biology and immature stages of Milichiella argyrogaster (Diptera, Milichiidae). – Ent. Probl. 26: 85-92)
fig. of eye, Algeria (Séguy 1934: 28. Diptères (Brachycères) (Muscidae Acalyptraterae et Scatophagidae): 832pp+27pl.: 636 (as Milichiella argrogastra))
catalog, Slovakia (Gregor 1986: Celad: Milichiidae, ?. – In: Cepelák, Diptera Slovenska, II: Cyclorrhapha, 435pp: 184)
diagnosis, larvae in decaying wood, males flying in hot sunny weather near cut old Populus logs and females running on these, Russia; Austria (Hennig 1937: Milichiidae et Carnidae, pp. 1-91. – In: Lindner, Die Fliegen der palaearktischen Region 61a: 26)
syn. to M. argyrogastra (Becker 1907: Die Dipteren-Gruppe Milichiinae. – Annls hist.-nat. Mus. natn. hung. 5: 507-550: 511 (as Lobioptera tiefi2))
Slovakia (Rohácek, J. & Gregor 1984: Nové nebo faunisticky zajímaví nálazy druhu celedi Milichiidae a Carnidae (Diptera) z Ceskoslovenska. – Cas. slezsk. Muz. Opave (A) 33: 17-21: 17)
Milichiella tiefi3: Becker 1905: Katalog der paläarktischen Dipteren 4: 237-240: 237 (as Milichiella tiefi3)
Milichiella tiefi3: Aldrich 1931: New acalyptrate Diptera from the Pacific and Oriental Regions. – Proc. Hawaii. ent. Soc. 7: 395-399: 398 (as Milichiella tiefi3)
on and swarming near stack of logs of old beech branches (Mik 1887: Über Dipteren. I. Drei neue österreichische Dipteren. – Verh. zool.-bot. Ges. Wien 37: 173-180,Taf.IV: 178 (as Lobioptera tiefii))
Russia: North European territory (Stackelberg 1958: Acalyptrata, part 1, pp. 103-191. – In: List of Diptera of the Leningrad Region. III: 186)
checklist, Slovakia (Gregor 1987: Milichiidae, p. 262. – In: Jezek, Enumeratio insectorum Bohemoslovakiae - Check list of Czechoslovak insects II (Diptera). 18: 262)