KEY TO THE SUBSPECIES

1Fruits globose, 3 cm diameter; pericarp 5-8 mm thick. — N Moluccas.
1'Fruist globose, less than 2.5 cm diameter, or (broadly) ellipsoid; pericarp 5 mm thick or less.
2Fruits ellipsoid, 3.5 cm long; fruiting pedicel 2 by 2 mm. — N Moluccas, Bacan.
2'Fruits subglobose or broadly ellipsoid, 1.8-2.3(-2.5) cm long; fruiting pedicel 3-5 mm.
3Fruits 2 cm long; pericarp (2-)3 mm thick. Lateral nerves on lower leaf surface not much raised but usually distinct; [midrib flat or raised above.] — W and E New Guinea main island (incl. Bird's Head and islands in the Geelvink Baai).
3'Fruits slightly larger (longer), 2.3(-2.5) cm long; pericarp (2-)3-5 mm thick. Lateral nerves on lower leaf surface faint.
4Leaves glossy on drying at both surfaces; midrib raised above. Fruits brightly rusty pubescent. Older twigs blackish, strongly finely flaking. — Central southern New Guinea.
4'Leaves rather dull on drying; midrib ± flat or but slightly raised above. Fruits chocolate or dull brown pubescent. Older twigs grey-brown, striate or slightly cracked, but not flaky. — Louisiade Archipelago.