REGIONAL KEY: BORNEO

1Stipules connate, 6-25 cm long
1'Stipules free, usually less than 6 cm long, on opening shoots, in some species up to 8 cm long
2Internodes conspicuously different in length, the proximal ones of a season’s growth long, subsequently shorter, the ultimate ones very short, sometimes with persistent stipules forming terminal buds
2'Internodes not conspicuously different in length
3Apex of the lamina caudate
3'Apex of the lamina acuminate, mostly shortly so
4Epidermis of petiole flaking off, at the uppermost or also at the basal part
4'Epidermis of petiole persistent
5Peduncle 0-0.1 cm long; basal bracts persistent
5'Peduncle 0.1-0.3(-0.5) cm long; basal bracts caducous
6Figs pedunculate or sessile with a peduncle-like stipe
6'Figs sessile
7Figs sessile with a peduncle-like stipe (the basal bracts at the base of the stipe)
7'Figs pedunculate (the basal bracts at the apex of the peduncle)
8Apex of the peduncle widened into a rim (bearing the basal bracts inside)
8'Apex of the peduncle not widened into a rim
9Midrib of the lamina beneath laterally hairy, often with hairs concentrated in the axils of the lateral veins
9'Midrib of the lamina beneath with hairs evenly distributed or absent
10Fig receptacle 0.4-0.7(-1) cm diam. when dry, the peduncle 0.7-1.2 cm long, 2 upper ostiolar bracts visible, these imbricate; leafy twigs glabrous or sparsely and minutely white puberulous
10'Fig receptacle 0.8-1.2 cm diam. when dry, the peduncle 0.2-0.7 cm long, 3 upper ostiolar bracts visible, these unequal in size and hardly or not imbricate; leafy twigs with dark brown appressed hairs
11Lateral and smaller veins (and often also the apex of the midrib) invisible
11'Lateral veins visible, smaller veins varying from clearly visible to ± obscure
12Tertiary venation parallel to the lateral veins (such as in F. elastica); lateral veins departing in wide angles (towards 90°)
12'Tertiary venation partly to largely parallel to the lateral veins to reticulate towards the margin or reticulate to subscalariform towards the margin; lateral veins usually departing in more acute angles (about 60°)
13Fig receptacle ellipsoid, ovoid, or cylindrical; petioles and stipules usually 1.5-3 cm long
13'Fig receptacle subglobose, or if distinctly longer than wide, then the petioles and/or stipules usually up to 1.5 cm long
14Lamina mostly up to 10 cm, rarely up to 15 cm long
14'Lamina mostly longer than 10 cm, up to c. 20 or up to c. 30 cm
15Fig receptacle longer than wide (ellipsoid, obovoid, ovoid, or cylindrical)
15'Fig receptacle about as long as wide (subglobose)
16Ostiole closed, the 3 upper ostiolar bracts clearly imbricate
16'Ostiole ± open, the 3 upper ostiolar bracts not or partly imbricate
17Fig receptacle 1.5-2 cm diam. when dry; basal bracts 5-10 mm long
17'Fig receptacle 0.5-1(-1.2) cm diam. when dry; basal bracts 3-5 mm long
18Basal lateral veins up to 1/10-1/4 the length of the lamina
18'Basal lateral veins up to 1/4-1/3(-1/2) the length of the lamina
19Petiole 0.5-1(-1.5) cm long; base of the lamina rounded to obtuse, apex rounded to obtuse to short-acuminate; fig receptacle ovoid
19'Petiole 1.5-2.5(-3) cm long; base of the lamina cuneate to obtuse, apex acuminate; fig receptacle ellipsoid
20Stipules distinctly hairy; margin of the lamina not callose towards the base; apex of the lamina acuminate
20'Stipules glabrous (or sparsely and minutely puberulous); margin of the lamina usually callose towards the base; apex of the lamina rounded to short-acuminate
21Ostiole ± open, the 3 upper ostiolar bracts not or partly imbricate
21'Ostiole closed, the 3 upper ostiolar bracts clearly imbricate
22Basal lateral veins branched, 1/3-1/2 the length of the lamina; margin of the lamina not callose towards the base
22'Basal lateral veins unbranched, mostly up to 1/4 the length of the lamina, or if up to 1/3(-1/2), then the margin of the lamina ± callose towards the base
23Ostiole (sub)conical
23'Ostiole flat, slightly prominent, or slightly sunken
24Fig receptacle 0.4-0.7 cm diam. when dry; stipules 0.5-1(-1.2) cm long
24'Fig receptacle (0.5-)0.7-1.2 cm diam. when dry; stipules (0.5-)1-1.5(-2) cm long
25Basal bracts 3-8(-10) mm long; fig receptacle 0.7-1.3(-1.8) cm diam. when dry
25'Basal bracts 1-3 mm long; fig receptacle 0.3-0.6(-0.8) cm diam. when dry, basal lateral veins up to 1/4-1/2 the length of the lamina, and/or petioles, stipules, and basal bracts usually glabrous
26Basal lateral veins up to 1/10-1/4 the length of the lamina; petioles, stipules, and basal bracts minutely white puberulous; margin of the lamina not callose towards the base
26'Basal lateral veins up to 1/4-1/3(-1/2) the length of the lamina; petioles, stipules, and basal bracts usually glabrous; margin of the lamina ± callose towards the base
27Petioles or stipules usually 1-2 cm long; basal lateral veins usually 1/10-1/4 the length of the lamina
27'Petioles and stipules usually 0.5-1 cm long; basal lateral veins usually 1/4-1/2 the length of the lamina
28Basal bracts 2-3 mm long; apex of the lamina short-acuminate (with an obtuse acumen) to obtuse to subacute to rounded
28'Basal bracts 1.5-2 mm long; apex of the lamina usually acuminate to subcaudate
29Stipules and/or petioles usually 0.5-1 cm long; apex of the lamina rounded to obtuse or to subacute to short-acuminate (with obtuse acumen); basal bracts 0.5-4 mm long
29'Stipules and petioles usually 1-1.5 cm long, or up to 3 or 4 cm long, respectively; apex of the lamina acuminate, or if rounded to obtuse, then the basal bracts 4-8(-10) mm long
30Tertiary venation (nearly) invisible
30'Tertiary venation visible
31Fig receptacle with internal hairs; midrib of the lamina flat to slightly prominent or impressed above; acumen of the lamina obtuse
31'Fig receptacle without internal hairs; midrib of the lamina impressed above; acumen of the lamina usually acute
32Indumentum of leafy twigs, petioles, and stipules (usually) brownish; basal bracts c. 2 mm long
32'Indumentum of leafy twigs, petioles, and stipules absent or whitish, or if brownish, then the basal bracts 3-5(-10) mm long
33Apex of the lamina acuminate, the acumen usually acute; margin of the lamina not callose
33'Apex of the lamina rounded to short-acuminate, the acumen usually obtuse; margin of the lamina ± callose towards the base
34Fig receptacle longer than wide
34'Fig receptacle about as long as wide (or wider than high)
35Fig receptacle 2-3 cm diam. when dry
35'Fig receptacle 0.3-2 cm diam. when dry
36Basal bracts 10-20(-30) mm long; tertiary venation of the lamina (sub)scalariform
36'Basal bracts 0.5-10 mm long; tertiary venation reticulate to subscalariform or partly parallel to the lateral veins (towards the midrib)
37Basal lateral veins (1/4-)1/3-1/2 the length of the lamina; tertiary venation of the lamina reticulate to subscalariform
37'Basal lateral veins up to 1/4 the length of the lamina, or if up to 1/3, then the tertiary venation of the lamina partly parallel to the lateral veins (towards the midrib)
38Stipules (1.5-)2-4 cm long; basal lateral veins unbranched, usually without smaller lateral veins below the (main) ones; base of the lamina rounded to cuneate (rarely subcordate)
38'Stipules 1-1.5(-2) cm long; (main) basal lateral veins often branched, often 1-3 pairs of smaller lateral veins below the main ones; base of the lamina cordate to rounded
39Fig receptacle 1-2 cm diam. when dry
39'Fig receptacle 0.3-1 cm diam. when dry
40Indumentum (partly) set(ul)ose with irritating hairs
40'Indumentum not set(ul)ose
41Basal bracts 0.5-3 mm long
41'Basal bracts 3-12 mm long
42Stipules 1-1.5(-2) cm long; base of the lamina cordate to rounded
42'Stipules 2-3 cm long; base of the lamina rounded to obtuse
43Petiole 0.5-1(-1.5) cm long, 1-2 mm thick
43'Petiole 1-5.5 cm long, 2-3(-5) mm thick
44Basal bracts 3-5 mm long; fig receptacle 0.5-1.2 cm diam. when dry
44'Basal bracts 5-8 mm long; fig receptacle 1.5-2 cm diam. when dry
45Ostiole ± open, the 3 upper ostiolar bracts not or slightly imbricate
45'Ostiole closed, the 3 upper ostiolar bracts clearly imbricate
46Base of the lamina rounded to cordate (with a narrow sinus); lamina puberulous on the venation beneath and on the midrib above
46'Base of the lamina cuneate to rounded; lamina glabrous
47Ostiole closed, the 3 upper ostiolar bracts clearly imbricate
47'Ostiole ± open, the 3 upper ostiolar bracts not or partly imbricate
48Basal bracts semicircular to suborbicular, imbricate, the apex rounded; areoles of lamina usually distinct beneath because of prominent veinlets
48'Basal bracts ovate, not (or only basally) imbricate, the apex obtuse; areoles of lamina obscure beneath
49Basal lateral veins branched; lateral veins (3-)4-6 pairs
49'Basal lateral veins (usually) unbranched; lateral veins 6-12 pairs
50Fig receptacle ellipsoid; petiole usually 1-2 cm long
50'Fig receptacle ovoid; petiole usually 0.5-1 cm long
51Fig receptacle 1-2 cm diam. when dry
51'Fig receptacle 0.3-1 cm diam. when dry
52Indumentum on various parts (as leafy twigs and midrib of the lamina beneath) brown floccose-tomentose
52'Indumentum not floccose-tomentose
53Apex of the fig receptacle concave, the ostiole sunken; stipules 2-6 cm long, subpersistent
53'Apex of the fig receptacle convex to submammillate, the ostiole ± prominent; stipules 1-2 cm long, mostly caducous
54Basal bracts 0.5-3 mm long
54'Basal bracts (3-)4-10(-18) mm long
55Lamina on the midrib above and the midrib and lateral veins beneath white puberulous; base of the lamina cordate to rounded
55'Lamina above and beneath glabrous or sometimes minutely puberulous on the midrib beneath; base of the lamina cuneate to rounded
56Midrib (at least the lower part) of lamina impressed above; base of lamina cordate to rounded
56'Midrib of lamina slightly prominent to flat; base of lamina usually cuneate to rounded
57Basal bracts semicircular to suborbicular with a rounded apex, imbricate, covering 1/3-2/3 of the receptacle; areoles of lamina usually distinct beneath because of prominent veinlets
57'Basal bracts (broadly) ovate, not (or only basally) imbricate, with an obtuse to acute or to rounded apex, covering up to 1/3 (or 1/2) of the receptacle
58Apex of the lamina acuminate, usually with an acute acumen; basal bracts ovate, not or only imbricate at the base; areoles of lamina obscure beneath
58'Apex of the lamina rounded to obtuse, or if acuminate, then with an obtuse acumen; basal bracts semicircular to broadly ovate, ± imbricate
59Base of the lamina and the lower part of the margin not (or hardly) callose; smaller veins of the lamina, even the reticulum ± clearly visible beneath; basal bracts usually covering 1/2 or more of the fig receptacle
59'Base of the lamina and (at least) the lower part of the margin ± distinctly callose; smaller veins of the lamina usually not clearly visible to obscure; basal bracts covering up to 1/2 of the fig receptacle
60Basal bracts 0.5-2.5 mm long
60'Basal bracts 3-8(-10) mm long
61Ostiole open, the 3 upper ostiolar bracts not or slightly imbricate; basal lateral veins distinct, up to 1/10-1/4 the length of the lamina
61'Ostiole closed, the 3 upper ostiolar bracts clearly imbricate; basal lateral veins indistinct, up to 1/20-1/10 the length of the lamina
62Indumentum on various parts (as stipules and midrib of the lamina beneath) brown floccose-tomentose
62'Indumentum not floccose-tomentose
63Leafy twigs (and often also on other parts) with abundant small dark brown appressed hairs; tertiary venation ± prominent beneath
63'Leafy twigs without such hairs or if present, then very sparse and inconspicuous; tertiary venation (almost) flat beneath
64Petiole 0.5-1 cm long; apex of the lamina rounded; tertiary venation of the lamina slightly prominent (rather inconspicuous)
64'Petiole (0.5-)1-2.5 cm long; apex of the lamina acuminate to rounded; tertiary venation prominent (and conspicuous)
65Ostiole closed, the 3 upper ostiolar bracts clearly imbricate
65'Ostiole ± open, the 3 upper ostiolar bracts not or partly imbricate
66Basal lateral veins up to 1/10-1/4 the length of the lamina
66'Basal lateral veins up to 1/4-1/2 the length of the lamina
67Fig receptacle 0.3-0.6 cm diam. when dry; basal bracts 1-2 mm long
67'Fig receptacle (0.5-)0.7-1.2 cm diam. when dry; basal bracts 3-5 mm long
68Lateral veins of the lamina 6-10 pairs; basal lateral veins unbranched, sometimes faintly branched
68'Lateral veins of the lamina (3-)4-6 pairs; basal lateral veins branched