Drynaria
Morphology
When the base fronds disintegrate, some species retain the naked rachises for a long time, sometimes with an empty mesh of finer veins.
Taxonomy
Drynaria is a distinct genus, best recognisable by the peculiar frond dimorphism between sterile base fronds and fertile foliage fronds, the distinctive leathery texture of the lamina, a distinctive venation pattern and the presence of nectaries near the bases of the pinnae. The last three characters are shared with Aglaomorpha. A curious character unique to Drynaria is the aborted apex, with a lateral pinna taking its place, giving a peculiar ‘lop-sided’ look to the apical part of most fronds.
Together with Aglaomorpha, Drynaria forms a distinct group, recognised as subfamily Drynarioideae by Crabbe, Jermy & Thomas (), and as tribe Drynarieae by Hennipman et al. (1990). Roos (1985) presents a cladistic analysis but does not propose a subdivision of Drynaria based on his analysis.
Together with Aglaomorpha, Drynaria forms a distinct group, recognised as subfamily Drynarioideae by Crabbe, Jermy & Thomas (), and as tribe Drynarieae by Hennipman et al. (1990). Roos (1985) presents a cladistic analysis but does not propose a subdivision of Drynaria based on his analysis.
Citation
1864 – In: Sp. Fil.: 93
Copel. 1947: Gen. Fil.: 204
Hook. & Bauer 1842: Gen. Fil.: t. 21
Alderw. 1917: Malayan Ferns: 415
Tind. 1961: Contr. NSW Nat. Herb.: 29
Ching 1940 – In: Sunyatsenia: 261
Hennipman et al. 1990 – In: Kramer & Green, Fam. & Genera Vasc. Pl. 1: 212
Holttum 1954 – In: Revis. Fl. Malaya: 182
Copel. 1905: Polypod. Philipp.: 135
Bedd. 1883: Handb. Ferns Brit. India: 338
Baker 1868: Syn. Fil.: 366
Hook. 1862: Gard. Ferns: pl. 1
Copel. 1929 – In: Univ. Calif. Publ. Bot.: 117
Copel. 1947: Gen. Fil.: 203
Alderw. 1908: Malayan Ferns: 696
J. Sm. 1875: Hist. Fil.: 107
Alderw. 1917: Malayan Ferns: 415
Fée 1850–1852: Gen. Filic.: 269
Copel. 1911 – In: Philipp. J. Sc. Bot.: 140
Morton 1970 – In: Taxon: 647
Alderw. 1908: Malayan Ferns: 697
Copel. 1960: Fern Fl. Philipp.: 496
Pichi Serm. 1977 – In: Webbia: 379
Backer & Posth. 1939: Varenfl. Java: 232
Roos 1985: Drynarioideae: 255
Alderw. 1908: Malayan Ferns: 679
Brownlie 1969 – In: Fl. Nouv. Caléd. et Dép.: 278
Bedd. 1863: Ferns S. India: 63
1917: Malayan Ferns: 415
Diels 1900 – In: Bot. Jahrb. Syst.: 207
Tardieu & C. Chr. 1941 – In: Fl. Indo-Chine: 516
J. Sm. 1866: Ferns Brit. & For.: 102