Drynaria
Morphology
When the base fronds disintegrate, some species retain the naked rachises for a long time, sometimes with an empty mesh of finer veins.
Taxonomy
Drynaria is a distinct genus, best recognisable by the peculiar frond dimorphism between sterile base fronds and fertile foliage fronds, the distinctive leathery texture of the lamina, a distinctive venation pattern and the presence of nectaries near the bases of the pinnae. The last three characters are shared with Aglaomorpha. A curious character unique to Drynaria is the aborted apex, with a lateral pinna taking its place, giving a peculiar ‘lop-sided’ look to the apical part of most fronds.
Together with Aglaomorpha, Drynaria forms a distinct group, recognised as subfamily Drynarioideae by Crabbe, Jermy & Thomas (), and as tribe Drynarieae by Hennipman et al. (1990). Roos (1985) presents a cladistic analysis but does not propose a subdivision of Drynaria based on his analysis.
Together with Aglaomorpha, Drynaria forms a distinct group, recognised as subfamily Drynarioideae by Crabbe, Jermy & Thomas (), and as tribe Drynarieae by Hennipman et al. (1990). Roos (1985) presents a cladistic analysis but does not propose a subdivision of Drynaria based on his analysis.
Citation
Alderw. 1908: Malayan Ferns. p 679
Copel. 1947: Gen. Fil. p 204
Diels 1900 – In: Bot. Jahrb. Syst. p 207
Holttum 1954 – In: Revis. Fl. Malaya. p 182
Roos 1985: Drynarioideae. p 255
Copel. 1947: Gen. Fil. p 203
Bedd. 1883: Handb. Ferns Brit. India. p 338
Tind. 1961: Contr. NSW Nat. Herb. p 29
Hennipman et al. 1990 – In: Kramer & Green, Fam. & Genera Vasc. Pl. 1. p 212
Copel. 1929 – In: Univ. Calif. Publ. Bot. p 117
Copel. 1960: Fern Fl. Philipp. p 496
Morton 1970 – In: Taxon. p 647
Copel. 1911 – In: Philipp. J. Sc. Bot. p 140
J. Sm. 1875: Hist. Fil. p 107
Baker 1868: Syn. Fil. p 366
Fée 1850–1852: Gen. Filic. p 269
Hook. & Bauer 1842: Gen. Fil. t. 21
1917: Malayan Ferns. p 415
1864 – In: Sp. Fil. p 93
Tardieu & C. Chr. 1941 – In: Fl. Indo-Chine. p 516
Alderw. 1917: Malayan Ferns. p 415
Brownlie 1969 – In: Fl. Nouv. Caléd. et Dép. p 278
Copel. 1905: Polypod. Philipp. p 135
Alderw. 1908: Malayan Ferns. p 696
Pichi Serm. 1977 – In: Webbia. p 379
Bedd. 1863: Ferns S. India. p 63
Alderw. 1917: Malayan Ferns. p 415
Hook. 1862: Gard. Ferns. pl. 1
J. Sm. 1866: Ferns Brit. & For. p 102
Backer & Posth. 1939: Varenfl. Java. p 232
Alderw. 1908: Malayan Ferns. p 697
Ching 1940 – In: Sunyatsenia. p 261