Asia-Temperate: Asia-Tropical:, Borneo present; Sulawesi (Sulawesi present); Vietnam (Vietnam present) East Asia: present Japan: present
East Asia: China, Japan, Vietnam. In Malesia: Borneo, Sulawesi.
This species is a hybrid between L. macrophyllus and, possibly, L. elliptica. It shows hybrid irregularity in blade outline. The plants have fewer sporangia and more paraphyses than the parent species. Few of the sporangia develop fully, and those that do fail to produce spores and do not dehisce in a normal manner