KEY TO THE SPECIES

1Leaves with conspicuously winged rachis — New Guinea
1'Leaf rachis unwinged, rarely weakly so
2Leaves opposite
2'Leaves spirally arranged
3Leaves pubescent, at least abaxially
3'Leaves glabrous
4Venation obscure
4'Costae and intercostae prominent
5Terminal leaflet present
5'Terminal leaflet absent; small ‘spike’ or scar ± present
6Inflorescences axillary
6'Inflorescences ramiflorous to cauliflorous
7Calyx splitting into lobes at anthesis
7'Calyx of discrete sepals
8Leaves up to 4-jugate; petals 5; fruit smooth
8'Leaves up to 8-jugate; petals 5; fruit cerebriform, tomentose
8''Leaves 5–8-jugate; petals 4; fruits smooth
9Indumentum of simple and stellate hairs; sepaloid bracteoles absent
9'Indumentum of simple hairs, sepaloid bracteoles present
10Leaves up to 3- (or 4-)jugate, treelet
10'Leaves with more leaflets
11Costae c. 16 on each side; inflorescence and fruit strigose-irritant; pachycaul treelet to 6 m
11'Costae c. 10–12 on each side; inflorescences not strigose
12Calyx 7–15 mm long
12'Calyx less than 4 mm long 13
13Leaves up to 8-jugate, leaflets strongly asymmetric; inflorescences ramiflorous; fruits silky hairy
13'Leaves 3–5-jugate; inflorescences ramiflorous to cauliflorous; fruits ± glabrous
14Staminal tube ± hairy. — Thailand to S Philippines
14'Staminal tube glabrous
15Inflorescences 3–8(–12) mm long; treelet. — New Guinea
15'Inflorescences much longer; trees.
16Inflorescences finely adpressed hairy. — E Malesia
16'Inflorescences densely strigose hairy. — C Malesia .
17Apical bud not spike-like, with young leaves like clenched fists
17'Apical bud spike-like or stiletto-shaped (sect. Cyrtochiton) .
18Leaves strictly paripinnate with terminal pair of leaflets .
18'Leaves not so
19Leaves 10–14-jugate
19'Leaves up to 7-jugate
20Costae c. 5–7 on each side, obscure; petals 4. — Malay Peninsula
20'Costae c. 6–9 on each side, obscure; petals 5. — Borneo .
20''Costae more, conspicuous. — Philippines, New Guinea
21Staminal tube unceolate (to subcylindrical); pollen shed in tetrads. — New Guinea
21'Staminal tube cylindrical, pollen grains shed singly
22Petals at least 8 mm long
22'Petals up to 5 mm long. — New Guinea
23Petals 4 — Philippines
23'Petals 5; capsule smooth. — New Guinea
23''Petals 5; capsule lenticellate-pustular. — New Guinea .
24Leaves 4–6-jugate; inflorescences 4–13 cm long; petals up to 2 mm long
24'Leaves 5–7-jugate; inflorescences to 35 cm long; petals c. 4.5 mm long
24''Leaves 3- or 4-jugate; inflorescences 4–9 cm long; petals at least 4 mm long.
25Leaves glabrous, venation markedly scalariform; petals 5 .
25'Leaves glabrous or not, venation different
26Leaflets narrowly elliptic (rheophyte). — Malay Peninsula
26'Leaflets not so
27Shoots and abaxial surface of leaflets densely reddish velutinous
27'Leaflets different
28Leaflets yellowish pilose abaxially; inflorescence spicate..
28'Leaflets and inflorescence different
29Costae up to 12 on each side
29'Costae (14) 15 on each side. — New Guinea
30Leaflets asymmetric at base, rarely subequal, shiny adaxially; bracteoles inconspicuous; calyx and corolla glabrous; infructescence ± branched; whole plant, or at least fruit, smelling of onions
30'Leaflets ± symmetric, not markedly shiny adaxially; bracteoles conspicuous; calyx and corolla pubescent; infructescence ± unbranched; whole plant not smelling of onions
31Bracts 5–10 by 1–2.5 mm; treelet with flagelliform inflorescences
31'Bracts up to 3.5 mm long; inflorescences to 20 cm long
32Petals up to 9 mm long
32'Petals more than 13 mm long
33Abaxial surface of leaflets brown-tomentose
33'Abaxial surface of leaflets glabrous to subglabrous
34Leaves 6–9-jugate; leaflets with 23–25 costae
34'Leaves 2–4-jugate; leaflets with 15–18 costae
35Intercostal venation scarcely distinguishable from costae
35'Costae distinct; intercostal venation often obscure
36Leaves paripinnate, 2–4(–5)-jugate, glabrous to puberulous
36'Leaves ± imparipinnate with ± alternate leaflets
37Petals 4–5 mm long; staminal tube crenulate. — New Guinea
37'Petals 8 mm long; staminal tube crenate to toothed
37''Petals 6 mm long; staminal tube truncate. — Borneo
38Leaflets to 30 by 10 cm; thyrses to 25 cm, with spreading branches
38'Leaflets smaller; thyrses subspiciform
39Leaflets minutely rugulose; costae 12–15(–18). — New Guinea
39'Leaflets not rugulose; costae 7–13
40Calyx c. 4 mm long. — West of Wallace’s Line
40'Calyx c. 1.5 mm long. — East of Wallace’s Line