| 1 | Leaves subverticillate, in a terminal whorl or clustered at intervals. Infructescences cauliflorous and/or axillary. |
| 1' | Leaves spaced, spiral. Infructescences often terminal, and/or axillary, rarely also cauliflorous. |
| 2 | Unbranched shrub-like treelet or slender tree. Papillae distinct on the lower leaf surface. Fruit obovoid or ellipsoid, 3-3½ cm long |
| 2' | Branched shrub or tree. Papillae indistinct or obscure on the lower leaf surface. Fruits unknown |
| 3 | Papillae indistinct or obscure on the lower leaf surface. |
| 3' | Papillae distinct on the lower leaf surface. |
| 4 | Fruits 2½-3½ cm Ø; hypocarp obconical 1-1½ cm Ø |
| 4' | Fruits wider, 4¼-5 cm Ø; hypocarp short-cupular, 3-3¼ cm Ø |
| 5 | Hypocarp discoid, short-cupular, or funnel-shaped. |
| 5' | Hypocarp pulvinate or obconical. |
| 6 | Fruits almost glabrous |
| 6' | Fruits pubescent, puberulous, or velutinous. |
| 7 | Leaf apex rounded or slightly apiculate; veins on lower leaf surface much elevated. Fruits velutinous; apex apiculate, sometimes concave at the top |
| 7' | Leaf apex acute or acuminate, sometimes obtuse, rarely slightly emarginate. |
| 8 | Fruits velutinous; apex concave |
| 8' | Fruits pubescent, glabrescent; apex obtuse |
| 9 | Papillae on lower leaf surface separated into small groups in the areolae. |
| 9' | Papillae rather uniform on the lower leaf surface. |
| 10 | Leaf apex obtuse, rounded, acute, sometimes shortly or abruptly acuminate. |
| 10' | Leaf apex shortly acuminate, slightly rostrate, or abruptly acuminate-rostrate. |
| 11 | Leaves with 5-10 pairs of nerves |
| 11' | Leaves with 22-32 pairs of nerves |
| 12 | Fruit apex shortly acuminate; hypocarp 1-1½ by 1-1½(-2) cm |
| 12' | Fruit apex acuminate-rostrate; hypocarp 2⅔ by 2⅓ cm |
