Asia-Tropical: Borneo present, S.W. Celebes present
Malesia: Borneo; S.W. Celebes (Brooks 16853, Maros).
One specimen from a shaded place at entrance to a cave has thinner pinnae more deeply lobed than above described, to 2.2 cm wide, costules 5 mm apart, pinna-lobes distinctly dentate at all vein-ends, veins often forked, only 1 pair anastomosing. A specimen from 1700 m on G. Mulu is small, with 5 pairs of free pinnae, largest 3.0 x 1.3 cm, hairs on lower surface of rachis and costae more than 0.5 mm long.