Unarmed, usually glabrous small trees, or erect or subscandent shrubs. Leaves simple, entire, or sometimes palmately 3-5-lobed, often with pellucid glands, without articulation between blade and petiole; Flowers sexually dimorphic. Petals 5, valvate. Stamens 5-8, extrorse. Ovary inferior, 4-8-celled; Fruit globose or ellipsoid, usually strongly ribbed when dry;
Asia-Temperate: Asia-Tropical:, Borneo present; Malaya (Peninsular Malaysia present) N. Sumatra: present tropical America: present
About 30 spp. in tropical America and eastern Asia south to Malesia (3 spp.): N. Sumatra, Malay Peninsula, Borneo. .
Distinguished from Hedera by the normally erect habit and the smooth (or only weakly ruminate) endosperm.
Harms - in E. & P., Nat. Pfl. Fam. 1900: 254
Philipson - in Bull. Br. Mus. Nat. Hist. Bot. 1951: 18
Clarke - in Fl. Br. Ind. 1879: 733
Harms - in E. & P., Nat. Pfl. Fam. 3. 1894: 40
Bth. - in B. & H., Gen. Pl. 1. 1865: 944
Merr. - in Brittonia. 1941: 129
Hui-Lin Li - in Sargentia. 1942: 38
Bth. - in B. & H., Gen. Pl. 1. 1865: 943
Hutch. - in Gen. Fl. Pl. 1967: 71
Stone - in Gard. Bull. Sing. 1977: 148
Philipson - in J. Bot. 1940: 116