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Trees or shrubs. Leaves paripinnate, 2-6-jugate; Inflorescences (supra-)axillary, thyrsoid, without or basally with 1 or 2 branches; Flowers unisexual, male and female ones only slightly different in perianth. Sepals 5, (almost) free, not or basally slightly imbricate, not petaloid, almost equal, appressed-hairy on both surfaces. Petals 5, usually oblique funnel-shaped, clawed, with 2 auricles at the apex of the claw. Stamens 8, exserted in the male flowers; Ovary 3-celled, hairy; Fruits sessile, 3-celled, obovoid to globular or top-shaped, loculicidal, circular or rounded to sharply deltoid in cross section, wall woody, up to 6.5 mm thick, outside glabrous to velutinous, inside tomentose, rarely (Solomon Islands, New Caledonia) glabrous. Seeds ellipsoid to obovoid, testa shiny black or rarely brown, sarcotesta covering only a small part of the seed, 2-lobed or shield-like.


Asia-Tropical, Australasia, New Caledonia present, New Hebrides present, Pacific: Fiji (Fiji present); Samoa (Samoa present); Tonga (Tonga present), Solomon Islands present, most of West Malesia absent
About 20 species, in Malesia (except most of West Malesia), Australia, Solomon Islands, New Hebrides, New Caledonia, Fiji, Samoa, and Tonga.


Radlk. 1985 – In: Fl. Austral.: 69
Radlk. 1920 – In: Bot. Jahrb.: 300
Radlk. 1933 – In: Engl., Pflanzenr. 98: 1258
Radlk. 1879 – In: Sitzungsber. Math.-Phys. CI. Königl. Bayer. Akad. Wiss. München: 502, 600
S.T. Reynolds 1985 – In: Austrobaileya: 154
Adema 1992 – In: Blumea: 541
Miq. 1859 – In: Fl. Ind. Bat.: 565
Backer & Bakh. f. 1965 – In: Fl. Java: 140
Koord. & Valeton 1903 – In: Bijdr. Booms. Java: 211