1 | Leaves opposite or clearly predominantly so; petals usually present |
1' | Leaves spiral or alternate; petals absent |
2 | Mangrove shrub or tree with stilt-roots and often pneumatophores; flowers with pair of small prophylls (bracteoles) fused to lower hypanthium |
2' | Woody lianas, shrubs or small trees, without stilt-roots and pneumatophores; flowers without prophylls (bracteoles) fused to lower hypanthium |
3 | Flowers in small dense spherical inflorescences; fruits densely packed into cone-like heads |
3' | Individual flowers clearly visible; fruits not in cone-like heads |
4 | Calyx lobes clearly demarcated; anthers dorsifixed and versatile |
4' | Calyx lobes scarcely developed; anthers adnate to filament |