Roupala suaveolens

<<<<Compound leaves>Blade>Base>Shape>Position

broader side of blade facing terminal leaflet1

1. 004-004-003-001-001

<<<<Simple leaves>Blade>Margins>Margin type>Serration pair number

1-3(-6) pairs2

2. 003-002-006-001-001

<<<Compound leaves>Blade>Base>Shape

strongly asymmetrical, , acute to obtuse, at times shortest side of blade acute, longer side obtuse3

3. 004-004-003-001

<<<Compound leaves>Blade>Margins>Margin type

entire4

4. 004-004-005-001

<<<Compound leaves>Blade>Secondary veins>Secondary vein pair number

5-8 pairs5

5. 004-004-006-001

<<<Compound leaves>Petiolule terminal leaflets>Base>Shape

obtuse to acute6

6. 004-006-003-001

<<<Compound leaves>Petiolule terminal leaflets>Margins>Margin type

entire7

7. 004-006-005-001

<<<Compound leaves>Petiolule terminal leaflets>Secondary veins>Secondary vein pair number

6-10 pairs8

8. 004-006-006-001

<<<Flowers>Filaments>Anthers>Dimensions

1.7-2.6 x 0.3-0.4 mm9

9. 007-002-003-001

<<<Fruits>Apex>Hairs>Colour

yellow-brown velutinous, sometimes slowly glabrescent10

10. 009-003-002-001

<<<Habit>Juvenile branches>Hairs>Colour

yellow-brown to red-brown, velutinous to long-tomentose, becoming short-tomentose and fading to grey11

11. 001-003-001-001

<<<Leaves>Blade>Hairs>Colour

drying brown, matt or shiny, densely yellow-brown or rufous-velutinous above and beneath when young, gradually glabrescent, hairs persisting on base of midvein above, and more generally beneath, fading to grey12

12. 002-001-002-001

<<<Simple leaves>Blade>Apex>Shape

sometimes acute, obtuse or acuminate13

13. 003-002-005-001

<<<Simple leaves>Blade>Base>Shape

obtuse- or rounded-decurrent, cordate or truncate, symmetrical, folded14

14. 003-002-004-001

<<<Simple leaves>Blade>Margins>Margin type

entire, or serrate when at transitional stage between simple and compound leaves with of serrations, not revolute15

15. 003-002-006-001

<<<Simple leaves>Petiole>Hairs>Colour

yellow-brown velutinous totomentose, fading to grey16

16. 003-001-003-001

<<<Simple leaves>Veins>Secondary veins>Secondary vein pair number

5-8(-12) pairs17

17. 003-003-002-001

<<Compound leaves>Blade>Apex

apex attenuate,18

18. 004-004-004

<<Compound leaves>Blade>Base

base ,19

19. 004-004-003

<<Compound leaves>Blade>Dimensions

4-11.5 x 1.5-4.2 cm20

20. 004-004-001

<<Compound leaves>Blade>Length relative to width

(length:breadth 2-3(-4.5):1)21

21. 004-004-002

<<Compound leaves>Blade>Margins

margin ,22

22. 004-004-005

<<Compound leaves>Blade>Secondary veins

secondary veins ;23

23. 004-004-006

<<Compound leaves>Dimensions>Length

13-26 cm long (including 2.5-8.5 cm long petiole)24

24. 004-001-001

<<Compound leaves>Dimensions>Width

7-19 cm broad25

25. 004-001-002

<<Compound leaves>Leaflets>Leaflet pair number

1-4 pairs26

26. 004-002-001

<<Compound leaves>Petiolule lateral leaflets>Length

0-0.8 mm long27

27. 004-003-001

<<Compound leaves>Petiolule terminal leaflets>Apex

apex attenuate,28

28. 004-006-004

<<Compound leaves>Petiolule terminal leaflets>Base

base ,29

29. 004-006-003

<<Compound leaves>Petiolule terminal leaflets>Dimensions

7.5-14 x 3.2-8 cm30

30. 004-006-001

<<Compound leaves>Petiolule terminal leaflets>Length

0-3 cm long31

31. 004-005-001

<<Compound leaves>Petiolule terminal leaflets>Length relative to width

(length:breadth 1.5-3.5:1)32

32. 004-006-002

<<Compound leaves>Petiolule terminal leaflets>Margins

margin ,33

33. 004-006-005

<<Compound leaves>Petiolule terminal leaflets>Secondary veins

secondary veins ;34

34. 004-006-006

<<Flower buds>Apex>Width

1.4-2 mm broad35

35. 006-001-001

<<Flower buds>Midlength>Width

0.7-1.2 mm broad at midlength36

36. 006-002-001

<<Flowers>Filaments>Anthers

anthers ;37

37. 007-002-003

<<Flowers>Filaments>Length

0.3-1 mm long38

38. 007-002-001

<<Flowers>Filaments>Relative distance

attached to tepals 5.5-9 mm from base39

39. 007-002-002

<<Flowers>Glands>Length

0.3-0.6 mm long40

40. 007-003-001

<<Flowers>Glands>Lobes

lobes in contact at base;41

41. 007-003-002

<<Flowers>Ovary hairs>Relative distance

extending (1-)1.5-2(-2.5) mm from base of ovary42

42. 007-004-001

<<Fruits>Apex>Constriction

not constricted to constricting for 2.5-3.5 mm43

43. 009-003-001

<<Fruits>Apex>Hairs

, if so, light brown-grey beneath44

44. 009-003-002

<<Fruits>Base>Constriction

constricted for (1-)3-7 mm45

45. 009-002-001

<<Fruits>Seeds>Dimensions

1.3-1.4 x 0.6-0.9 cm46

46. 009-004-001

<<Habit>Bark>Colour

dark grey to red-grey or brown47

47. 001-004-001

<<Habit>Lenticels>Colour

orange to brown48

48. 001-005-002

<<Habit>Lenticels>Lenticel number

few49

49. 001-005-001

<<Habit>Shrub habit>Height

1.5-2.5 m tall50

50. 001-002-001

<<Habit>Tree habit>Height

3-10(-20) m51

51. 001-001-001

<<Inflorescences>Bracts>Dimensions

1-2 x 0.6-1 mm52

52. 005-007-001

<<Inflorescences>Bracts>Hairs

densely velutinous to tomentose outside, glabrous within53

53. 005-007-002

<<Inflorescences>Flowers>Length

to 1mm long54

54. 005-008-001

<<Inflorescences>Hairs>Colour

pale brown, yellow-brown to rufous55

55. 005-004-001

<<Inflorescences>Pedicels>Length

1.5-5 mm long56

56. 005-009-001

<<Inflorescences>Peduncle>Dimensions

1-3.5(-6) cm x 1.2-2 mm57

57. 005-005-001

<<Infructescences>Fruit pedicels>Dimensions

(3-)4-6 x 1-1.5 mm58

58. 008-003-001

<<Infructescences>Fruit pedicels>Hairs

tomentose59

59. 008-003-002

<<Leaves>Blade>Texture

coriaceous60

60. 002-001-001

<<Simple leaves>Blade>Apex

apex predominantly attenuate, ,61

61. 003-002-005

<<Simple leaves>Blade>Base

base ,62

62. 003-002-004

<<Simple leaves>Blade>Dimensions

6-21 x 3-12 cm63

63. 003-002-002

<<Simple leaves>Blade>Length relative to width

(length:breadth 1.1-2.5:1)64

64. 003-002-003

<<Simple leaves>Blade>Margins

margin ;65

65. 003-002-006

<<Simple leaves>Blade>Shape

ovate, also wide- to narrow-ovate, or narrow-oblong to oblong66

66. 003-002-001

<<Simple leaves>Petiole>Hairs

, rarely becoming completely glabrous67

67. 003-001-003

<<Simple leaves>Petiole>Length

1-6 cm long68

68. 003-001-001

<<Simple leaves>Petiole>Width

1.5-4 mm broad at midlength69

69. 003-001-002

<<Simple leaves>Veins>Midveins

midvein reaching apex,70

70. 003-003-001

<<Simple leaves>Veins>Secondary veins

secondary veins .71

71. 003-003-002

<Compound leaves>Blade

blade lateral leaflet ,72

72. 004-004

<Compound leaves>Dimensions

, ;73

73. 004-001

<Compound leaves>Juvenile plants

compound leaves of juvenile plants with considerably longer and narrower leaves, long-attenuate at apex and long-cuneate at base.74

74. 004-007

<Compound leaves>Leaflets

leaflets ;75

75. 004-002

<Compound leaves>Petiolule lateral leaflets

petiolules lateral leaflets ;76

76. 004-003

<Compound leaves>Petiolule terminal leaflets

blade terminal leaflet ,77 petiolule terminal leaflet ,78

77. 004-006, 78. 004-005

<Flower buds>Apex

at apex79

79. 006-001

<Flowers>Filaments

filaments , ,80

80. 007-002

<Flowers>Glands

hypogynous glands ,81

81. 007-003

<Flowers>Length

7-11 mm long82

82. 007-001

<Flowers>Ovary hairs

ovary hairs , covering entire ovary, long, weak, straight, ferrugineous to rufous, also with sericeous hairs, dense, bushing outwards.83

83. 007-004

<Fruits>Apex

apex , sharp with persistence of style, straight or curved so apex perpendicular to unsutured side, , marginal vein protruding around periphery;84

84. 009-003

<Fruits>Base

base ,85

85. 009-002

<Fruits>Dimensions

1.8-3.2 x 0.9-1.5 cm86

86. 009-001

<Fruits>Seeds

seeds .87

87. 009-004

<Habit>Bark

bark ;88

88. 001-004

<Habit>Juvenile branches

young branches , persistent;89

89. 001-003

<Habit>Lenticels

lenticels , inconspicuous, .90

90. 001-005

<Habit>Shrub habit

shrub91

91. 001-002

<Habit>Tree habit

Tree92

92. 001-001

<Inflorescences>Bracts

common bracts , ;93

93. 005-007

<Inflorescences>Dimensions

7-28 x 1.8-2.8 cm94

94. 005-003

<Inflorescences>Flowers

flower-pair axis ;95

95. 005-008

<Inflorescences>Hairs

densely , velutinous or tomentose96

96. 005-004

<Inflorescences>Inflorescence number per leaf axil

rarely 297

97. 005-002

<Inflorescences>Pedicels

pedicels .98

98. 005-009

<Inflorescences>Peduncle

peduncle ;99

99. 005-005

<Inflorescences>Position

axillary100

100. 005-001

<Inflorescences>Sterile bracts

sterile bracts few, persistent;101

101. 005-006

<Infructescences>Fruit pedicels

fruit pedicel , .102

102. 008-003

<Infructescences>Hairs

densely tomentose103

103. 008-002

<Infructescences>Length

9-21(-34) cm long104

104. 008-001

<Leaves>Blade

blade ,105

105. 002-001

<Simple leaves>Blade

blade predominantly , ,106

106. 003-002

<Simple leaves>Petiole

petiole x , terete, somewhat canaliculate towards blade, ;107

107. 003-001

<Simple leaves>Veins

venation raised and ± conspicuous on both sides, eucamptodromous to semicraspedodromous,108

108. 003-003

Compound leaves

Compound leaves:109

109. 004

Description

Tree 3-10(-20) m, or shrub 1.5-2.5 m tall; young branches yellow-brown to red-brown, velutinous to long-tomentose, becoming short-tomentose and fading to grey, persistent; bark dark grey to red-grey or brown; lenticels few, inconspicuous, orange to brown. Leaves simple, lobed and compound, blade coriaceous, drying brown, matt or shiny, densely yellow-brown or rufous-velutinous above and beneath when young, gradually glabrescent, hairs persisting on base of midvein above, and more generally beneath, fading to greySimple leaf: petiole 1-6 cm long x 1.5-4 mm broad at midlength, terete, somewhat canaliculate towards blade, yellow-brown velutinous totomentose, fading to grey, rarely becoming completely glabrous; blade predominantly ovate, also wide- to narrow-ovate, or narrow-oblong to oblong, 6-21 x 3-12 cm(length:breadth 1.1-2.5:1), base obtuse- or rounded-decurrent, cordate or truncate, symmetrical, folded, apex predominantly attenuate, sometimes acute, obtuse or acuminate, margin entire, or serrate when at transitional stage between simple and compound leaves with 1-3(-6) pairs of serrations, not revolute; venation raised and ± conspicuous on both sides, eucamptodromous to semicraspedodromous, midvein reaching apex, secondary veins 5-8(-12) pairs. Compound leaves: 13-26 cm long (including 2.5-8.5 cm long petiole), 7-19 cm broad; leaflets 1-4 pairs; petiolules lateral leaflets 0-0.8 mm long; blade lateral leaflet 4-11.5 x 1.5-4.2 cm(length:breadth 2-3(-4.5):1), base strongly asymmetrical, broader side of blade facing terminal leaflet, acute to obtuse, at times shortest side of blade acute, longer side obtuse, apex attenuate, margin entire, secondary veins 5-8 pairs; petiolule terminal leaflet 0-3 cm long, blade terminal leaflet 7.5-14 x 3.2-8 cm(length:breadth 1.5-3.5:1), base obtuse to acute, apex attenuate, margin entire, secondary veins 6-10 pairs; compound leaves of juvenile plants with considerably longer and narrower leaves, long-attenuate at apex and long-cuneate at base. Inflorescence axillary, unbranched, rarely 2 arising in same leaf axil, 7-28 x 1.8-2.8 cm, densely pale brown, yellow-brown to rufous, velutinous or tomentose; peduncle 1-3.5(-6) cm x 1.2-2 mm; sterile bracts few, persistent; common bracts 1-2 x 0.6-1 mm, densely velutinous to tomentose outside, glabrous within; flower-pair axis to 1mm long; pedicels 1.5-5 mm long. Flower buds 1.4-2 mm broad at apex, 0.7-1.2 mm broad at midlengthFlowers 7-11 mm long; filaments 0.3-1 mm long, attached to tepals 5.5-9 mm from base, anthers 1.7-2.6 x 0.3-0.4 mm; hypogynous glands 0.3-0.6 mm long, lobes in contact at base; ovary hairs extending (1-)1.5-2(-2.5) mm from base of ovary, covering entire ovary, long, weak, straight, ferrugineous to rufous, also with sericeous hairs, dense, bushing outwards. Infructescence 9-21(-34) cm long, densely tomentose; fruit pedicel (3-)4-6 x 1-1.5 mm, tomentose. Fruit 1.8-3.2 x 0.9-1.5 cm, both sutured and unsutured side curved to similar degrees, or sutured side slightly more curved, base constricted for (1-)3-7 mm, apex not constricted to constricting for 2.5-3.5 mm, sharp with persistence of style, straight or curved so apex perpendicular to unsutured side, yellow-brown velutinous, sometimes slowly glabrescent, if so, light brown-grey beneath, marginal vein protruding around periphery; seeds 1.3-1.4 x 0.6-0.9 cm.

Distribution

Bolivar present, Guyana present, Southern America: Brazil North (Amazonas present), Venezuela where rivers drain northwards into the Orinoco system present

Venezuela where rivers drain northwards into the Orinoco system (Amazonas and Bolívar) , and Guyana; (GU: 6; SU: 1).

Flower buds

Flower buds ,110

110. 006

Flowers

Flowers ;111

111. 007

Fruits

Fruit , both sutured and unsutured side curved to similar degrees, or sutured side slightly more curved,112

112. 009

Habit

, or ;113

113. 001

Individuals Association

Guyana, Mazaruni-Potaro Distr., Mt. Roraima, Arabupu, Forest Dept. 2814 Guyana, Rupununi Distr., Kamoa R., Toucan Mt., Jansen-Jacobs et al. 1628

Inflorescences

Inflorescence , unbranched, arising in same leaf axil, ,;114

114. 005

Infructescences

Infructescence , ;115

115. 008

Leaves

Leaves simple, lobed and compound,116

116. 002

Notes

Roupala schomburgkii was originally differentiated from R. suaveolens by the shortness of the petiole, differing shapes of leaves and leaf bases. However, these are inconsistant characters which vary independently of each other in different specimens. Roupala griotii is clearly synonymous with R. suaveolens.
Although the locality on the type specimen label of Roupala suaveolens is difficult to interpret from the handwriting, according to , this species was seen in the vicinity of Mt. Amboina, Pirocaima and Camana, now in Roraima State, Brazil. The type of R. schomburgkii was from the locality of Mt. Roraima which was ascended from the Brazilian side, hence the change in type localities from Guyana to Brazil.

Phenology

Flowering from , fruiting predominantly from .

Simple leaves

Simple leaf:117

117. 003

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