Roupala suaveolens

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Roupala suaveolens

Description

Tree 3-10(-20) m, or shrub 1.5-2.5 m tall; young branches yellow-brown to red-brown, velutinous to long-tomentose, becoming short-tomentose and fading to grey, persistent; bark dark grey to red-grey or brown; lenticels few, inconspicuous, orange to brown. Leaves simple, lobed and compound, blade coriaceous, drying brown, matt or shiny, densely yellow-brown or rufous-velutinous above and beneath when young, gradually glabrescent, hairs persisting on base of midvein above, and more generally beneath, fading to greySimple leaf: petiole 1-6 cm long x 1.5-4 mm broad at midlength, terete, somewhat canaliculate towards blade, yellow-brown velutinous totomentose, fading to grey, rarely becoming completely glabrous; blade predominantly ovate, also wide- to narrow-ovate, or narrow-oblong to oblong, 6-21 x 3-12 cm(length:breadth 1.1-2.5:1), base obtuse- or rounded-decurrent, cordate or truncate, symmetrical, folded, apex predominantly attenuate, sometimes acute, obtuse or acuminate, margin entire, or serrate when at transitional stage between simple and compound leaves with 1-3(-6) pairs of serrations, not revolute; venation raised and ± conspicuous on both sides, eucamptodromous to semicraspedodromous, midvein reaching apex, secondary veins 5-8(-12) pairs. Compound leaves: 13-26 cm long (including 2.5-8.5 cm long petiole), 7-19 cm broad; leaflets 1-4 pairs; petiolules lateral leaflets 0-0.8 mm long; blade lateral leaflet 4-11.5 x 1.5-4.2 cm(length:breadth 2-3(-4.5):1), base strongly asymmetrical, broader side of blade facing terminal leaflet, acute to obtuse, at times shortest side of blade acute, longer side obtuse, apex attenuate, margin entire, secondary veins 5-8 pairs; petiolule terminal leaflet 0-3 cm long, blade terminal leaflet 7.5-14 x 3.2-8 cm(length:breadth 1.5-3.5:1), base obtuse to acute, apex attenuate, margin entire, secondary veins 6-10 pairs; compound leaves of juvenile plants with considerably longer and narrower leaves, long-attenuate at apex and long-cuneate at base. Inflorescence axillary, unbranched, rarely 2 arising in same leaf axil, 7-28 x 1.8-2.8 cm, densely pale brown, yellow-brown to rufous, velutinous or tomentose; peduncle 1-3.5(-6) cm x 1.2-2 mm; sterile bracts few, persistent; common bracts 1-2 x 0.6-1 mm, densely velutinous to tomentose outside, glabrous within; flower-pair axis to 1mm long; pedicels 1.5-5 mm long. Flower buds 1.4-2 mm broad at apex, 0.7-1.2 mm broad at midlengthFlowers 7-11 mm long; filaments 0.3-1 mm long, attached to tepals 5.5-9 mm from base, anthers 1.7-2.6 x 0.3-0.4 mm; hypogynous glands 0.3-0.6 mm long, lobes in contact at base; ovary hairs extending (1-)1.5-2(-2.5) mm from base of ovary, covering entire ovary, long, weak, straight, ferrugineous to rufous, also with sericeous hairs, dense, bushing outwards. Infructescence 9-21(-34) cm long, densely tomentose; fruit pedicel (3-)4-6 x 1-1.5 mm, tomentose. Fruit 1.8-3.2 x 0.9-1.5 cm, both sutured and unsutured side curved to similar degrees, or sutured side slightly more curved, base constricted for (1-)3-7 mm, apex not constricted to constricting for 2.5-3.5 mm, sharp with persistence of style, straight or curved so apex perpendicular to unsutured side, yellow-brown velutinous, sometimes slowly glabrescent, if so, light brown-grey beneath, marginal vein protruding around periphery; seeds 1.3-1.4 x 0.6-0.9 cm.

Distribution

Bolivar present, Guyana present, Southern America: Brazil North (Amazonas present), Venezuela where rivers drain northwards into the Orinoco system present
Venezuela where rivers drain northwards into the Orinoco system (Amazonas and Bolívar) , and Guyana; (GU: 6; SU: 1).

Phenology

Flowering from , fruiting predominantly from .

Notes

Roupala schomburgkii was originally differentiated from R. suaveolens by the shortness of the petiole, differing shapes of leaves and leaf bases. However, these are inconsistant characters which vary independently of each other in different specimens. Roupala griotii is clearly synonymous with R. suaveolens.
Although the locality on the type specimen label of Roupala suaveolens is difficult to interpret from the handwriting, according to , this species was seen in the vicinity of Mt. Amboina, Pirocaima and Camana, now in Roraima State, Brazil. The type of R. schomburgkii was from the locality of Mt. Roraima which was ascended from the Brazilian side, hence the change in type localities from Guyana to Brazil.