Herbs, monoecious (or dioecious). Leaves spirally arranged, rosulate; stipules semi-amplexicaul, free, persistent; blade entire or lobed to parted, pinnately to subpalmately veined. Inflorescences in the leaf axils, bisexual or unisexual, pedunculate; receptacle discoid to cup-shaped, with (sub)marginal bracts. Staminate flowers numerous, superficial; tepals 2(-3), free; stamens 2; pistillode absent. Pistillate flowers numerous, embedded in the receptacle; perianth tubular; ovary free, stigmas 2, subulate. Fruit free, a dehiscent drupelet, the white, fleshy exocarp ejecting the endocarp body; seed small, with endosperm, cotyledons flat and equal.