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Herbs, monoecious (or dioecious). Leaves spirally arranged, rosulate; stipules semi-amplexicaul, free, persistent; blade entire or lobed to parted, pinnately to subpalmately veined. Inflorescences in the leaf axils, bisexual or unisexual, pedunculate; receptacle discoid to cup-shaped, with (sub)marginal bracts. Staminate flowers numerous, superficial; tepals 2(-3), free; stamens 2; pistillode absent. Pistillate flowers numerous, embedded in the receptacle; perianth tubular; ovary free, stigmas 2, subulate. Fruit free, a dehiscent drupelet, the white, fleshy exocarp ejecting the endocarp body; seed small, with endosperm, cotyledons flat and equal.


Africa present, Asia present, Guianas present, Neotropics present
In Africa and adjacent parts of Asia with approx. 60 species and in the Neotropics with approx. 45 species. In the Guianas 1, possibly 3 species.