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Terrestrial, caulescent, erect herbs, without modified stems. Stems unbranched. Leaves opposite, nearly equal in a pair, venation pinnate, foliar nectaries absent. Flowers axillary, 1 to few in umbellate-cymose inflorescences; short pedunculate; bracteoles small; long pedicellate. Calyx lobes 5, free; corolla yellowish, tubular, inflated ventrally, limb 5-lobed; stamens included, filaments basally connate, anthers free or coherent in 2 pairs, dehiscing by longitudinal slits, thecae parallel; staminode present, small; disc of 2 or 4 opposite separate glands, or a double connate dorsal gland; ovary superior, stigma capitate or 2-lobed. Fruit a fleshy, indehiscent, black berry.


Bolivar present, Guayana Highlands of Venezuela present, Guianas present, Southern America: Bolivia (Bolivia present); Brazil North (Amazonas present); Venezuela (Venezuela present), western Guyana present
An Andean genus of 8 or more species, in mixed wet or cloud forest, known from Bolivia to coastal Venezuela, and in the Guayana Highlands of Venezuela (BolĂ­var, Amazonas), neighbouring Brazil, and western Guyana; 1 species in the Guianas.


Chromosome number n=9 (Skog 1984).