Primary tabs



Shrubs or trees with dichotomous branching; Leaves opposite; Inflorescence a cyme or solitary flower, lax; Sepals in most species with a distinct connate tube and free lobes; Stamens subsessile; Fruit of paired follicles, usually somewhat united at least at base. Seeds covered in a fleshy aril.


Africa present, Asia-Tropical, Australasia: Queensland (Queensland present)
The genus has 12 species of which 7 are in Africa and 5 in Malesia. One of the Malesian species also occurs in Australia (Queensland).


This account is partly adapted from Leeuwenberg (1985) rather than completely reworked because I agree with the species delimitation he used. It has been updated to include new collections made since 1985.


Miq. 1864 – In: Ann. Mus. Bot. Lugduno-Batavi: 316
Thouars 2001 – In: PROSEA: 582
Miq. 1857 – In: Fl. Ned. Ind.: 417
D.J. Middleton 2004 – In: Tree Fl. Sabah & Sarawak: 59
Leeuwenb. 1985 – In: Agric. Univ. Wageningen Pap.: 9
Backer & Bakh.f. 1965 – In: Fl. Java: 229
Pichon 1947 – In: Bull. Mus. Hist. Nat. (Paris): 409