Pisonia macranthocarpa

<<<Habit>Stems>Spines>Length

0.3-1.1 cm long1

1. 001-003-001-002

<<<Habit>Stems>Spines>Position

axillary2

2. 001-003-001-001

<<Anthocarps>Glands>Row number

53

3. 006-007-001

<<Anthocarps>Glands>Series

often uniseriate, sometimes 2-seriate4

4. 006-007-002

<<Female perianth>Style>Length

1.2 mm5

5. 005-003-001

<<Habit>Bark>Colour

reddish6

6. 001-002-002

<<Habit>Bark>Texture

smooth7

7. 001-002-001

<<Habit>Stems>Hairs

densely pubescent, becoming glabrescent with age8

8. 001-003-002

<<Habit>Stems>Spines

often armed with , stout, straight to slightly curved, spines9

9. 001-003-001

<<Inflorescences>Bracteoles>Length

1-1.4 mm long10

10. 003-005-002

<<Inflorescences>Bracteoles>Shape

oblong11

11. 003-005-001

<<Inflorescences>Female inflorescences>Length

3.6-10 cm long12

12. 003-003-001

<<Inflorescences>Male inflorescences>Length

1.5-3.5 cm long13

13. 003-002-001

<<Inflorescences>Pedicels>Hairs

short, viscid-pubescent14

14. 003-006-001

<<Inflorescences>Peduncle>Length

1-5 cm15

15. 003-004-001

<<Petiole>Blade>Dimensions

11 x 6 cm16

16. 002-002-003

<<Petiole>Blade>Hairs

glabrous to sparsely or densely puberulent on upper surface, glabrous or shortly villous on lower surface17

17. 002-002-004

<<Petiole>Blade>Shape

ovate, elliptical, obovate or suborbicular18

18. 002-002-002

<<Petiole>Blade>Texture

subcoriaceous19

19. 002-002-001

<Anthocarps>Angle or rib number

5-angled and -ribbed20

20. 006-005

<Anthocarps>Colour

green21

21. 006-002

<Anthocarps>Dimensions

16-20 x 8-9 mm22

22. 006-004

<Anthocarps>Glands

with rows of stipitate glands, glands in each row23

23. 006-007

<Anthocarps>Hairs

densely puberulent24

24. 006-006

<Anthocarps>Shape

obconical or clavate25

25. 006-003

<Anthocarps>Texture

coriaceous26

26. 006-001

<Female perianth>Length

1.6-3 mm long27

27. 005-002

<Female perianth>Shape

tubular to more or less urceolate28

28. 005-001

<Female perianth>Style

style .29

29. 005-003

<Habit>Bark

bark , ;30

30. 001-002

<Habit>Height

to 6 m or more31

31. 001-001

<Habit>Stems

stems , .32

32. 001-003

<Inflorescences>Bracteoles

bracteoles , ;33

33. 003-005

<Inflorescences>Female inflorescences

female34

34. 003-003

<Inflorescences>Male inflorescences

male35

35. 003-002

<Inflorescences>Pedicels

pedicels .36

36. 003-006

<Inflorescences>Peduncle

peduncle ;37

37. 003-004

<Inflorescences>Position

axillary38

38. 003-001

<Male perianth>Colour

greenish-yellow39

39. 004-001

<Male perianth>Dimensions

2-4 x 2 mm40

40. 004-003

<Male perianth>Hairs

minutely puberulent-papillate41

41. 004-004

<Male perianth>Shape

broadly campanulate42

42. 004-002

<Male perianth>Stamens

stamens exserted.43

43. 004-005

<Petiole>Blade

blade , , , .44

44. 002-002

<Petiole>Length

to 3 cm long45

45. 002-001

Anthocarps

Anthocarp on elongated pedicel, dry, , , , , , , .46

46. 006

Description

Scrambling shrub or liana with numerous arcuate branches, climbing to 6 m or more; bark smooth, reddish; stems often armed with axillary, stout, straight to slightly curved, 0.3-1.1 cm long spines, densely pubescent, becoming glabrescent with age. Petiole to 3 cm long; blade subcoriaceous, ovate, elliptical, obovate or suborbicular, 11 x 6 cm, glabrous to sparsely or densely puberulent on upper surface, glabrous or shortly villous on lower surface. Inflorescence seemingly axillary, usually borne at apex of very reduced short shoots, male 1.5-3.5 cm long, female 3.6-10 cm long; peduncle 1-5 cm; bracteoles oblong, 1-1.4 mm long; pedicels short, viscid-pubescent. Male perianth greenish-yellow, broadly campanulate, 2-4 x 2 mm, minutely puberulent-papillate; stamens exserted. Female perianth tubular to more or less urceolate, 1.6-3 mm long; style 1.2 mm. Anthocarp on elongated pedicel, dry, coriaceous, green, obconical or clavate, 16-20 x 8-9 mm, 5-angled and -ribbed, densely puberulent, with 5 rows of stipitate glands, glands in each row often uniseriate, sometimes 2-seriate.

Distribution

Central and South America present, Guyana present

Central and South America, including Guyana where occurring in thickets on wooded, elevated "islands" in savanna vegetation; 20 non-Guianan collections studied; 1 collection from a single locality in Guyana, was examined: Rupununi Distr., Chaakoitou, near Mountain Point, just S of Kanuku Mts., (male flowers in bud), Maas et al. 4117 (K, U).

Female perianth

Female perianth , ;47

47. 005

Habit

Scrambling shrub or liana with numerous arcuate branches, climbing ;48

48. 001

Inflorescences

Inflorescence seemingly , usually borne at apex of very reduced short shoots, , ;49

49. 003

Male perianth

Male perianth , , , ;50

50. 004

Petiole

Petiole ;51

51. 002

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