Piper adenandrum

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Piper adenandrum

Content

<<<Petiole>Blade>Base>Differential distance

0-0.2 cm1
1. 002-002-005-001

<<Inflorescences>Peduncle>Length

0.5-1.1 cm long2
2. 003-002-001

<<Inflorescences>Spikes>Length

1-5 cm long3
3. 003-003-001

<<Petiole>Blade>Apex

apex long-acuminate,4
4. 002-002-004

<<Petiole>Blade>Base

base unequally attached to petiole difference , acute or obtusish, glabrous or pubescent above, (densely) crisp-pubescent or villous beneath;5
5. 002-002-005

<<Petiole>Blade>Dimensions

8-16.5 x 3-6(-7) cm6
6. 002-002-002

<<Petiole>Blade>Margins

margin ciliate,7
7. 002-002-003

<<Petiole>Blade>Shape

lance-elliptic or elliptic-ovate8
8. 002-002-001

<Fruits>Diameter

2.5-3(-4) mm in diam.9
9. 004-001

<Fruits>Stigma

stigmas 3, sessile or fruits somewhat stylose.10
10. 004-002

<Habit>Height

1.5 m tall11
11. 001-001

<Habit>Length

to 0.5 mm long12
12. 001-002

<Habit>Upper internodes

upper internodes densely glandular.13
13. 001-003

<Inflorescences>Anthers

anthers laterally dehiscent.14
14. 003-006

<Inflorescences>Floral bracts

floral bracts densely marginally fringed;15
15. 003-005

<Inflorescences>Growth form

pendent16
16. 003-001

<Inflorescences>Peduncle

peduncle , sometimes reddish tinged, crisp-pubescent or villous;17
17. 003-002

<Inflorescences>Rachises

rachis glabrous;18
18. 003-004

<Inflorescences>Spikes

spike , green to pale or dark yellow, apiculate;19
19. 003-003

<Petiole>Blade

blade not conspicuously glandular-dotted, , ,20
20. 002-002

<Petiole>Length

0.2-0.5(-1) cm long21
21. 002-001

<Petiole>Veins

pinnately veined, secondary veins 5-6 per side, originating from lower 3/4 of primary vein.22
22. 002-003

Common Name

Boni (French Guiana): ampukuwiwii English (Guyana): tona tonakeng English (Suriname): katun uwi

Description

Shrub or treelet, 1.5 m tall, pubescent or villous, hairs to 0.5 mm long; upper internodes densely glandular. Petiole 0.2-0.5(-1) cm long, vaginate to middle; blade not conspicuously glandular-dotted, lance-elliptic or elliptic-ovate, 8-16.5 x 3-6(-7) cm, margin ciliate, apex long-acuminate, base unequally attached to petiole difference 0-0.2 cm, acute or obtusish, glabrous or pubescent above, (densely) crisp-pubescent or villous beneath; pinnately veined, secondary veins 5-6 per side, originating from lower 3/4 of primary vein. Inflorescence pendent; peduncle 0.5-1.1 cm long, sometimes reddish tinged, crisp-pubescent or villous; spike 1-5 cm long, green to pale or dark yellow, apiculate; rachis glabrous; floral bracts densely marginally fringed; anthers laterally dehiscent. Fruits separate at maturity, depressed globose, 2.5-3(-4) mm in diam., verruculose; stigmas 3, sessile or fruits somewhat stylose.

Distribution

Guianas present, Southern America: Venezuela (Venezuela present)
Venezuela and the Guianas; 81 collections studied (GU: 14; SU: 5; FG: 62).

Fruits

Fruits separate at maturity, depressed globose, , verruculose;23
23. 004

Habit

Shrub or treelet, , pubescent or villous, hairs ;24
24. 001

Individuals Association

Kaw Mts., Feuillet 2240 Guyana, Upper Takutu — Upper Essequibo region, Gunn’s, Clarke 7059 Suriname, Lely Mts., Lindeman & Stoffers et al. 834 Saül area, foot Mt. Galbao, de Granville et al. 8398 Guyana, Acarai Mts., Clarke 2947 Suriname, Gran Santi, Sauvain 448

Inflorescences

Inflorescence ;25
25. 003

Notes

Piper adenandrum and P. avellanum had been considered conspecific by Kramer & Görts-van Rijn (1968: 418). There are, however, several differentiating characters: P. adenandrum has long-acuminate blades, indument of spreading, to 0.5 mm long hairs and distinctly separate fruits, 2.5-3(-4) mm in diam., whereas P. avellanum blades are (short) acuminate or acute, the hairs are crisp, 0.2 mm long, and the fruits 2 mm in diam., not distinctly separate.

Petiole

Petiole , vaginate to middle;26
26. 002